Colombia 2013

a non-profit small group tour organised and led by Adrian Hoskins

Trip report

The view from our lodge at Tatama national park – Adrian Hoskins

Things don’t always go according to plan in South America, but it all adds to the sense of adventure! This was our first trip to Colombia, and when we arrived we found the country was in the middle of a national strike which caused disruption and road closures on major routes. Fortunately with the aid of our excellent local guides Dora and Hernan, and our safe reliable driver Mauricio we were able to rearrange the schedule to avoid the road blocks, and a fantastic time was had by everyone!

Delicious food, comfortable accommodation, stunning scenery, gorgeous butterflies, great company and wonderful Colombian hospitality made this a very special tour!

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Marpesia%20corinna%201508 002b small - Butterflies Store

After spending a night at Medellin we flew to Pereira to avoid the road blocks, and then had a short drive to our first butterfly site Otun Quimbaya. Our lodge there was located amidst cloudforest at an elevation of 1900m, and most of our butterfly-watching and photography took place on the little-used forest road that stretches from the reserve entrance at 1800m up to about 2300m where the road ends. We also explored the riverbanks, and walked along the narrow trails that led from the lodge into the surrounding forest, where we saw the clearwing Satyr Pseudohaetera hypaesia, and a good selection of Ithomiine glasswings.

The latter are fascinating to watch, fluttering secretively with deep slow wing beats, and occasionally settling on leaves to pose for photographs. Downhill from the lodge we explored a grassy track at the edge of the river, which produced the stunning zebra hairstreak Arawacus leucogyna, several very attractive red and metallic blue-black Altinote species, and a huge red skipper Yanguna cosyra.

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Cloudforest at Otun Quimbaya – Adrian Hoskins

Following two full days of immersive butterfly photography at Otun, we embarked on a lengthy journey to our next destination: Tatama National Park. Nestled at an elevation of 1350 meters, our lodge was situated on the fringes of this enchanting and picturesque cloud forest. Our explorations primarily unfolded along the rugged stony road, meandering upwards from the lodge to nearly 3000 meters.

Tatama lies within the Choco ecological zone, renowned for receiving some of the highest rainfall in South America. Yet, it also stands as one of the continent’s most biodiverse butterfly havens. Although intermittent cloud cover and light drizzles deterred us from venturing to the highest elevations, we found ample opportunities for discovery between 1350-1800 meters, where conditions remained relatively warm and dry.

Some among us opted for a leisurely uphill ride in vehicles, followed by a slow descent on foot back to the lodge. Others chose to trek all the way to 1800 meters and back—an exhausting but immensely rewarding endeavor. Along the way, we encountered a multitude of butterfly species, including Callicore, Diaethria, Memphis, Heliconius, Dismorphia, Catasticta, Perisama, Adelpha, Doxocopa, Mesosemia, Catonephele, Epiphile, Hypanartia, Marpesia, and Corades. Among these sightings were the majestic Owl butterfly Caligo oedipus and three resplendent blue giant Morpho species.

The highlights of our journey were undoubtedly the breathtaking pink Fountainea nobilis, the charming Necyria bellona zaneta—a diminutive metalmark adorned with metallic blue bands and crimson spots against a jet-black backdrop—and the magnificent Epiphile eriopis, boasting striking black, red, and white hues as it basked gracefully with outstretched wings on the forest track.

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Cecropia tree at dawn, Tatama national park – Adrian Hoskins

Although our tour was primarily focused on butterflies, we often found ourselves distracted by the many other wonders of Tatama – stunning scenery, hummingbirds galore, beautiful orchids, moths of every shape and pattern imaginable, gorgeous dragonflies, and magnificent bugs!

Unidentified bug, Tatama national park – Adrian Hoskins

After reveling in six delightful days at Tatama, our journey led us to Rio Claro, where we spent three days at a nature resort nestled along the riverbanks at an elevation of 350 meters. While butterfly abundance in this locale was relatively modest, the diversity of species proved to be abundant. We encountered and captured photographs of numerous species not found in the higher elevations we had previously visited. Among these treasures were several exquisite metalmarks, including pristine specimens of Sarota chrysus, Ancyluris jurgensenii, Caria mantinea, Caria rhacotis, Lasaia agesilas, and Rhetus periander.

Our next stop took us to the Arrierito Piha Reserve in Antioquoia. Nestled along a tranquil forest road at an elevation of 1700 meters, the cozy lodge provided a wonderful retreat, particularly enchanting for hummingbird enthusiasts. At times, up to 40 of these mesmerizing creatures graced a favored small tree in the lodge garden.

Along the roadside, we encountered a plethora of fabulous butterflies, including the dead-leaf mimic Consul panariste, the stunning metallic blue-banded Emperors Doxocopa cyane and Doxoxopa laurentia cherubina, the charming white-banded Oressinoma typhla, and the blue-tailed metalmark Rhetus dysonii.

Traversing the trails through the cloudforest interior yielded further delights, such as the imposing and remarkably attractive Ithomiine Elzunia humboldt and the gracefully elegant longwing Heliconius clysonymus. Another noteworthy discovery was the white-banded ringlet Splendeuptychia ackeryi, a species that made its scientific debut in 2009, adding a special touch of novelty to our expedition.

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Hypanartia%20lethe%20001b small - Butterflies Store

Following our two-day sojourn at the Piha reserve, we embarked on a short drive to Santa Fe de Antioquoia, where a lengthy roadside stop rewarded us with sightings of the vividly colored longwing Dryadula phaetusa, various species of Urbanus long-tailed skippers, and a multitude of attractive hairstreak species. Another nearby site yielded sightings of Monarchs Danaus plexippus and the magnificent banded hairstreak Panthiades bathildis.

Our final day was spent in the company of two friends, Juan and Martin, residents of Medellin who are deeply knowledgeable and passionate about butterfly photography. They graciously guided us to a seemingly unremarkable patch of relict cloudforest on the outskirts of Medellin, situated at an elevation of 1400 meters. To our astonishment, this humble locale proved to be one of the most fruitful sites of our entire journey! A liberal application of liquidized fish bait onto foliage swiftly attracted swarms of butterflies, including the metallic blue Heliconius erato chestertonii and several alluring Adelpha species. Yet, the butterflies that captivated us most were the distinctive Eyemark Skipper Cyclosemia leppa and the stunningly attractive furry-legged Jewelmark Anteros kupris.

In the afternoon, we ventured to another nearby site at an elevation of 1800 meters, where we were treated to sightings of the pretty green hairstreak Cyanophrys longula, numerous glasswing and tiger-patterned Ithomiines, and a variety of pronophiline Satyrs, among which was a Forsterinaria species previously unknown to science.

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Pueblito de San Jose, near Medellin – Adrian Hoskins

SPECIES LIST

Species recorded in Colombia in August 2013 – list updated 28 Dec 2013

In total we saw about 430 species on the tour, of which 421 were photographed by the participants Adrian Hoskins, Peter Chadd, Peter Bygate, Les Catchick and Chris Orpin.

Most of us photographed between 200-300 species. The list which follows was compiled by Adrian Hoskins after meticulously checking all the images taken by the participants. Resources consulted included the 7 indispensable volumes of d’Abrera’s ‘Butterflies of the Neotropical region’, Keith Willmott’s ‘The Genus Adelpha’, and the Butterflies of America (Neotropical) website.

I’d like to express my thanks also to Andrew Neild, Tomasz Pyrcz, Jean Francois le Crom, Luis Miguel Constantino and Blanca Huertas for their valued assistance in determining some of the more difficult species.

Hundreds of photographs of the species seen on the tour can now be seen in the Amazon & Andes section, where you will also find detailed articles about the biology of each species.

Hesperiidae	Achylodes	busirus	heros
Hesperiidae	Achylodes	pallida
Hesperiidae	Anastrus	meliboea	meliboea
Hesperiidae	Anisochoria	pedaliodina	polysticta
Hesperiidae	Anthoptus	epictetus	epictetus
Hesperiidae	Antigonus	nearchus	nearchus
Hesperiidae	Apaustus	gracilis	gracilis
Hesperiidae	Aroma	aroma
Hesperiidae	Astraptes	anaphus
Hesperiidae	Astraptes	chiriquensis
Hesperiidae	Astraptes	fulgerator
Hesperiidae	Autochton	longipennis
Hesperiidae	Autochton	neis
Hesperiidae	Autochton	zarex
Hesperiidae	Bolla	dorsolaciniae
Hesperiidae	Bolla	giselus
Hesperiidae	Camptopleura	auxo	giselus
Hesperiidae	Camptopleura	impressus
Hesperiidae	Camptopleura	theramenes
Hesperiidae	Carrhenes	bamba	bamba
Hesperiidae	Carrhenes	canescens	canescens
Hesperiidae	Celaenorrhinus	eligius
Hesperiidae	Celaenorrhinus	suthina
Hesperiidae	Chalypyge	zereda	rufipectus
Hesperiidae	Cogia	calchas
Hesperiidae	Corticea	mendica
Hesperiidae	Cycloglypha	enega
Hesperiidae	Cycloglypha	thrasibulus	thrasibulus
Hesperiidae	Cycloglypha	tisias	tisias
Hesperiidae	Cyclosemia	leppa
Hesperiidae	Cymaenes	trebius
Hesperiidae	Cymaenes	tripunctus	theogenis
Hesperiidae	Dalla	calima
Hesperiidae	Dalla	eryonas
Hesperiidae	Dalla	frater
Hesperiidae	Dalla	plancus	plancus
Hesperiidae	Enosis	achelous
Hesperiidae	Epargyreus	aspina
Hesperiidae	Falga	jeconia
Hesperiidae	Gindanes	brebisson	panaetius
Hesperiidae	Gorgythion	begga	begga
Hesperiidae	Helias	cama
Hesperiidae	Helias	godmani
Hesperiidae	Helias	phalaenoides
Hesperiidae	Heliopetes	alana
Hesperiidae	Heliopetes	arsalte	arsalte
Hesperiidae	Heliopetes	laviana	laviana
Hesperiidae	Hylephila	phyleus	phyleus
Hesperiidae	Lento	ludo
Hesperiidae	Mictris	crispus	caerula
Hesperiidae	Milanion	hemes
Hesperiidae	Mylon	lassia
Hesperiidae	Mylon	zephus
Hesperiidae	Nisoniades	hecale
Hesperiidae	Noctuana	haematospila
Hesperiidae	Noctuana	noctua
Hesperiidae	Nyctelius	nyctelius	nyctelius
Hesperiidae	Ouleus	fridericus	fridericus
Hesperiidae	Panoquina	evadnes	evadnes
Hesperiidae	Pellicia	santana
Hesperiidae	Penicula	cristina
Hesperiidae	Phocides	pigmalion	pigmalion
Hesperiidae	Polythrix	hirtius
Hesperiidae	Pompeius	pompeius
Hesperiidae	Potamanaxas	andaemon
Hesperiidae	Potamanaxas	laoma	trigga
Hesperiidae	Potamanaxas	latrea	caliadne
Hesperiidae	Potamanaxas	melicertes
Hesperiidae	Potamanaxas	quira
Hesperiidae	Pyrgus	adepta
Hesperiidae	Pyrgus	orcus
Hesperiidae	Pyrrhopyge	evansi	evansi
Hesperiidae	Pyrrhopyge	papius	papius
Hesperiidae	Pythonides	herennius	herennius
Hesperiidae	Quadrus	lugubris	ophia
Hesperiidae	Remella	remus
Hesperiidae	Saliana	hewitsoni
Hesperiidae	Saliana	salius	salius
Hesperiidae	Saliana	severus
Hesperiidae	Spioniades	abbreviata	anta
Hesperiidae	Timochreon	satyrus	satyrus
Hesperiidae	Urbanus	albimargo	albimargo
Hesperiidae	Urbanus	esta
Hesperiidae	Urbanus	proteus
Hesperiidae	Urbanus	simplicius	simplicius
Hesperiidae	Urbanus	teleus	teleus
Hesperiidae	Vettius	coryna	catargyra
Hesperiidae	Vettius	marcus
Hesperiidae	Xenophanes	tryxus	tryxus
Hesperiidae	Yanguna	cosyra
Hesperiidae	Zenis	jebus	beckeri
Hesperiidae	Zera	hosta
Lycaenidae	Arawacus	aetolus	aetolus
Lycaenidae	Arawacus	leucogyna
Lycaenidae	Arawacus	lincoides
Lycaenidae	Brangas	felderi
Lycaenidae	Calycopis	calus
Lycaenidae	Calycopis	cerata
Lycaenidae	Calycopis	demonassa
Lycaenidae	Contrafacia	imma
Lycaenidae	Cupido	comyntas	texana
Lycaenidae	Cyanophrys	longula
Lycaenidae	Erora	subflorens
Lycaenidae	Gargina	emessa
Lycaenidae	Hemiargus	hanno
Lycaenidae	Ipidecla	schausi
Lycaenidae	Laothus	gibberosa
Lycaenidae	Leptotes	bathyllos
Lycaenidae	Micandra	platyptera
Lycaenidae	Ministrymon	azia	azia
Lycaenidae	Ministrymon	una
Lycaenidae	Mithras	colombiensis
Lycaenidae	Nicolaea	heraldica
Lycaenidae	Nicolaea	pyxis
Lycaenidae	Ocaria	aholiba
Lycaenidae	Ostrinotes	keila
Lycaenidae	Panthiades	bathildis
Lycaenidae	Pseudolycaena	marsyas
Lycaenidae	Rekoa	meton
Lycaenidae	Strymon	mulucha
Lycaenidae	Strymon	serapio
Lycaenidae	Symbiopsis	tanais
Lycaenidae	Theclopsis	demea
Lycaenidae	Thereus	orasus
Lycaenidae	Theritas	augustinula
Lycaenidae	Theritas	hemon
Lycaenidae	Theritas	mavors
Lycaenidae	Tmolus	echion	echiolus
Lycaenidae	Zizula	cyna
Nymphalidae	Actinote	anteas	anteas
Nymphalidae	Actinote	pellenea	limbata
Nymphalidae	Adelpha	alala	negra
Nymphalidae	Adelpha	barnesia	leucas
Nymphalidae	Adelpha	boeotia	boeotia
Nymphalidae	Adelpha	cytherea	daguana
Nymphalidae	Adelpha	ethelda	eponina
Nymphalidae	Adelpha	justina	justina
Nymphalidae	Adelpha	lamasi
Nymphalidae	Adelpha	leucopthalma	leucopthalma
Nymphalidae	Adelpha	levona
Nymphalidae	Adelpha	lycorias	wallisii
Nymphalidae	Adelpha	melona	deborah
Nymphalidae	Adelpha	rothschildi
Nymphalidae	Adelpha	serpa	celerio
Nymphalidae	Aeria	eurimedia	latistriga
Nymphalidae	Altinote	neleus	neleus
Nymphalidae	Altinote	ozomene
Nymphalidae	Altinote	stratonice	stratonice
Nymphalidae	Amphidecta	calliomma
Nymphalidae	Anartia	amathea	amathea
Nymphalidae	Anartia	jatrophae	jatrophae
Nymphalidae	Anthanassa	acesas	castianira
Nymphalidae	Anthanassa	drusilla	drusilla
Nymphalidae	Archaeoprepona	demophon	muson
Nymphalidae	Archaeoprepona	meander	megabates
Nymphalidae	Caligo	illioneus	illioneus
Nymphalidae	Caligo	oedipus	oedipus
Nymphalidae	Callicore	atacama	manova
Nymphalidae	Callicore	lyca
Nymphalidae	Callicore	pitheas
Nymphalidae	Callicore	tolima	denina
Nymphalidae	Castilia	castilla	castilla
Nymphalidae	Castilia	eranites
Nymphalidae	Catonephele	chromis	godmani
Nymphalidae	Catonephele	numilia	esite
Nymphalidae	Catonephele	nyctimus
Nymphalidae	Ceratinia	tutia	transversa
Nymphalidae	Chlosyne	narva	bonpland
Nymphalidae	Chlosyne	theona	hondana
Nymphalidae	Cissia	labe
Nymphalidae	Colobura	annulata	annulata
Nymphalidae	Colobura	dirce	dirce
Nymphalidae	Consul	fabius
Nymphalidae	Consul	panariste	ludmilla
Nymphalidae	Corades	enyo	almo
Nymphalidae	Corades	pannonia	ploas
Nymphalidae	Dagon	pusilla
Nymphalidae	Danaus	eresimus	montezuma
Nymphalidae	Danaus	plexippus
Nymphalidae	Diaethria	clymena	marchalii
Nymphalidae	Diaethria	euclides	phlogea
Nymphalidae	Dione	juno	andicola
Nymphalidae	Dione	moneta	butleri
Nymphalidae	Dircenna	dero
Nymphalidae	Doxocopa	cyane	mexicana
Nymphalidae	Doxocopa	laurentia	cherubina
Nymphalidae	Doxocopa	pavon
Nymphalidae	Doxocopa	zunilda	felderi
Nymphalidae	Dryadula	phaetusa
Nymphalidae	Dryas	iulia	iulia
Nymphalidae	Dynamine	chryseis
Nymphalidae	Dynamine	paulina	thalassina
Nymphalidae	Dynamine	postverta	postverta
Nymphalidae	Dynamine	sosthenes	sosthenes
Nymphalidae	Ectima	erycinoides
Nymphalidae	Elzunia	humboldt	humboldt
Nymphalidae	Epiphile	chrysites	chrysites
Nymphalidae	Epiphile	dilecta
Nymphalidae	Epiphile	eriopis	eriopis
Nymphalidae	Eresia	carme
Nymphalidae	Eresia	clio	clio
Nymphalidae	Eresia	datis	manto
Nymphalidae	Eresia	emerantia	emerantia
Nymphalidae	Eresia	ithomioides	poecilina
Nymphalidae	Eresia	levina
Nymphalidae	Eresia	phaedima
Nymphalidae	Eresia	polina	polina
Nymphalidae	Eretris	calisto	calisto
Nymphalidae	Eretris	depresissima
Nymphalidae	Eretris	ocellifera	ocellifera
Nymphalidae	Eretris	oculata
Nymphalidae	Eunica	bechina	bechina
Nymphalidae	Eunica	carias	carias
Nymphalidae	Eunica	evelide	evelide
Nymphalidae	Eunica	norica	norica
Nymphalidae	Euptychia	jesia
Nymphalidae	Euptychoides	griphe
Nymphalidae	Euptychoides	laccine
Nymphalidae	Euptychoides	saturnus
Nymphalidae	Forsterinaria	inornata
Nymphalidae	Forsterinaria	neonympha
Nymphalidae	Forsterinaria	pallida	aurita
Nymphalidae	Forsterinaria	pseudinornata
Nymphalidae	Fountainea	nessus	nessus
Nymphalidae	Fountainea	nobilis	pacifica
Nymphalidae	Gnathotriche	exclamationis	exclamationis
Nymphalidae	Godyris	duillia
Nymphalidae	Godyris	zavaleta
Nymphalidae	Greta	andromica	andromica
Nymphalidae	Hamadryas	fornax
Nymphalidae	Hamadryas	laodamia
Nymphalidae	Heliconius	charithonia	charithonia
Nymphalidae	Heliconius	clysonymus	clysonymus
Nymphalidae	Heliconius	cydno	cydnides
Nymphalidae	Heliconius	doris	obscuris, viridis & dives
Nymphalidae	Heliconius	erato	chestertonii
Nymphalidae	Heliconius	hecale
Nymphalidae	Heliconius	melpomene	martinae
Nymphalidae	Hermeuptychia	cucullina
Nymphalidae	Hermeuptychia	harmonia
Nymphalidae	Hermeuptychia	hermes
Nymphalidae	Hermeuptychia	maimoune
Nymphalidae	Historis	odius	odius
Nymphalidae	Hypanartia	dione	dione
Nymphalidae	Hypanartia	lethe
Nymphalidae	Hypanartia	trimaculata	trimaculata
Nymphalidae	Hypna	clytemnestra
Nymphalidae	Hypoleria	lavinia	vanilia
Nymphalidae	Hypothyris	lycaste
Nymphalidae	Ithomia	avella	cesleria
Nymphalidae	Ithomia	hyala	sincerus
Nymphalidae	Janatella	fellula
Nymphalidae	Janatella	leucodesma
Nymphalidae	Junonia	evarete	zonalis
Nymphalidae	Lycorea	halia
Nymphalidae	Lymanopoda	albocincta	albocincta
Nymphalidae	Lymanopoda	obsoleta
Nymphalidae	Magneuptychia	libye
Nymphalidae	Magneuptychia	tiessa
Nymphalidae	Manataria	hercyna	daguana
Nymphalidae	Marpesia	chiron
Nymphalidae	Marpesia	corinna
Nymphalidae	Marpesia	furcula
Nymphalidae	Marpesia	marcella	marcella
Nymphalidae	Marpesia	petreus
Nymphalidae	Marpesia	zerynthia
Nymphalidae	Mechanitis	polymnia	veritabilis
Nymphalidae	Melinaea	idea
Nymphalidae	Memphis	acidalia	memphis
Nymphalidae	Memphis	arginussa
Nymphalidae	Memphis	laura	caucana
Nymphalidae	Memphis	lyceus
Nymphalidae	Memphis	moeris
Nymphalidae	Memphis	oenomais
Nymphalidae	Memphis	perenna	austrina
Nymphalidae	Memphis	xenocles
Nymphalidae	Mesotaenia	vaninka	doris
Nymphalidae	Morpho	deidamia	granadensis
Nymphalidae	Morpho	helenor	leontius
Nymphalidae	Napeogenes	peridia	hoppi
Nymphalidae	Nessaea	aglaura	aglaura
Nymphalidae	Nica	flavilla
Nymphalidae	Oleria	amalda	amalda
Nymphalidae	Oleria	makrena
Nymphalidae	Oleria	phenomoe
Nymphalidae	Oleria	santineza
Nymphalidae	Opsiphanes	cassina	chiriquensis
Nymphalidae	Opsiphanes	invirae	sticheli
Nymphalidae	Opsiphanes	quiteria	badius
Nymphalidae	Oressinoma	typhla	typhla
Nymphalidae	Oxeoschistus	simplex	fuscus
Nymphalidae	Panacea	prola
Nymphalidae	Parataygetis	lineata	lineata
Nymphalidae	Pareuptychia	hesionides
Nymphalidae	Pareuptychia	metaleuca
Nymphalidae	Pareuptychia	ocirrhoe	ocirrhoe
Nymphalidae	Patricia	dercyllidas	dercyllidas
Nymphalidae	Pedaliodes	dejecta
Nymphalidae	Pedaliodes	demarmelsi
Nymphalidae	Pedaliodes	phrasicla	phrasicla
Nymphalidae	Perisama	bomplandii	bomplandii
Nymphalidae	Perisama	humboldtii	humboldtii
Nymphalidae	Perisama	oppelii	oppelii
Nymphalidae	Perisama	typhania
Nymphalidae	Perisama	yeba	malvina
Nymphalidae	Philaethria	dido	dido
Nymphalidae	Pierella	helvina	helvina
Nymphalidae	Podotricha	judith	caucana
Nymphalidae	Pronophila	unifasciata	brennus
Nymphalidae	Pseudohaetera	hypaesia
Nymphalidae	Pseudohaetera	mimica
Nymphalidae	Pseudomaniola	phaselis	pholoe
Nymphalidae	Pseudoscada	timna	saturata
Nymphalidae	Pyrrhogyra	crameri	undine
Nymphalidae	Pyrrhogyra	edocla	edocla
Nymphalidae	Siproeta	epaphus
Nymphalidae	Siproeta	stelenes
Nymphalidae	Smyrna	blomfildia	datis
Nymphalidae	Taygetis	chrysogone
Nymphalidae	Taygetis	thamyra
Nymphalidae	Tegosa	claudina	claudina
Nymphalidae	Temenis	laothoe	colombiana
Nymphalidae	Tigridia	acesta	acesta
Nymphalidae	Tithorea	tarricina	parola
Nymphalidae	Vanessa	braziliensis
Nymphalidae	Ypthimoides	renata
Papilionidae	Battus	polydamas	polydamas
Papilionidae	Eurytides	colombus
Papilionidae	Heraclides	thoas	nealces
Papilionidae	Neographium	agesilaus	eimeri
Papilionidae	Parides	erithalion	erithalion
Pieridae	Anteos	maerula
Pieridae	Archonias	brassolis	semialba
Pieridae	Ascia	monuste	monuste
Pieridae	Catasticta	ctemene	strigosa
Pieridae	Catasticta	flisa	dilutior
Pieridae	Catasticta	hegemon	helle
Pieridae	Catasticta	prioneris	albescens
Pieridae	Catasticta	sisamnus
Pieridae	Dismorphia	crisia	foedora
Pieridae	Dismorphia	hippotas	diana
Pieridae	Dismorphia	lelex	valeria
Pieridae	Dismorphia	lua	idea
Pieridae	Dismorphia	medora	medora
Pieridae	Dismorphia	mirandola	discolora
Pieridae	Dismorphia	theucharila	theucharila
Pieridae	Dismorphia	zaela	zaela
Pieridae	Dismorphia	zathoe	zathoe
Pieridae	Enantia	lina	agatha
Pieridae	Eurema	albula	marginella
Pieridae	Eurema	arbela	gratiosa
Pieridae	Eurema	daira	lydia
Pieridae	Eurema	elathea	vitellina
Pieridae	Eurema	xantochlora	xantochlora
Pieridae	Hesperocharis	marchallii
Pieridae	Leodonta	tellane	intermedia
Pieridae	Leptophobia	aripa	aripa
Pieridae	Leptophobia	caesia	tenuicornis
Pieridae	Leptophobia	tovaria	maruga
Pieridae	Lieinix	nemesis	nemesis
Pieridae	Melete	leucanthe	leucanthe
Pieridae	Phoebis	neocypris	rurina
Pieridae	Phoebis	philea
Pieridae	Phoebis	sennae
Pieridae	Pseudopieris	nehemia	luisa
Pieridae	Pyrisitia	nise	stygma
Pieridae	Pyrisitia	venusta	venusta
Riodinidae	Amphiselenis	chama
Riodinidae	Ancyluris	inca	huascar
Riodinidae	Ancyluris	jurgensenii	atahualpa
Riodinidae	Anteros	cruentatus
Riodinidae	Anteros	formosus	theleia
Riodinidae	Anteros	kupris	kupris
Riodinidae	Argyrogrammana	caelestina
Riodinidae	Baeotis	macularia
Riodinidae	Baeotis	zonata
Riodinidae	Calephelis	laverna	laverna
Riodinidae	Caria	mantinea
Riodinidae	Caria	rhacotis	rhacotis
Riodinidae	Chamaelimas	briola
Riodinidae	Chalodetta	lypera
Riodinidae	Charis	anius	anius
Riodinidae	Crocozona	pheretima
Riodinidae	Detritivora	hermodora
Riodinidae	Emesis	cypria	paphia
Riodinidae	Emesis	glaucescens
Riodinidae	Emesis	lucinda
Riodinidae	Emesis	lupina
Riodinidae	Emesis	mandana	mandana
Riodinidae	Emesis	ocypore
Riodinidae	Emesis	tegula
Riodinidae	Emesis	tenedia
Riodinidae	Eurybia	lycisca
Riodinidae	Euselasia	bettina
Riodinidae	Euselasia	corduena	anadema
Riodinidae	Euselasia	eupatra
Riodinidae	Exoplisia	hypochalybe
Riodinidae	Lasaia	agesilas	agesilas
Riodinidae	Leucochimona	icare	polita
Riodinidae	Leucochimona	lagora
Riodinidae	Lyropteryx	apollonia
Riodinidae	Melanis	cinaron
Riodinidae	Melanis	electron	pronostriga
Riodinidae	Menander	pretus
Riodinidae	Mesosemia	mevania	mevania
Riodinidae	Mesosemia	minos	modica
Riodinidae	Mesosemia	pacifica
Riodinidae	Mesosemia	zonalis
Riodinidae	Napaea	theages	asteria
Riodinidae	Necyria	bellona	zaneta & westwoodi
Riodinidae	Notheme	erota	diadema
Riodinidae	Rhetus	dysonii	caligosus
Riodinidae	Rhetus	periander	laonome
Riodinidae	Sarota	chrysus
Riodinidae	Sarota	neglecta
Riodinidae	Siseme	alectryo	alectryo
Riodinidae	Siseme	aristoteles	sprucei
Riodinidae	Siseme	pallas	pallas
Riodinidae	Synargis	calyce
Riodinidae	Theope	pedias
Riodinidae	Thisbe	irenea

Trip report

SPECIES LIST

Butterflies of the Amazon & Andes

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