In August 2004, a non-profit small group trip organized and led by Adrian Hoskins took us on an enthralling journey through the diverse landscapes of West Malaysia and Borneo.
Our adventure began in Tapah on the Malayan peninsula, where we encountered the magnificent Rajah Brook’s Birdwing Trogonoptera brookiana in Kuala Woh Park. Stunning mud-puddling groups of these creatures, with as many as 30 individuals, adorned the landscape with their iridescent green and black wings outstretched. Edging down the riverbank, I found myself amidst these captivating creatures, experiencing absolute bliss.
While most other species at Kuala Woh were found in ones and twos, we encountered some notable finds including the long-tailed, bright orange hairstreak Eooxylides tharis, the gorgeous Euploea radamanthus, Appias lyncida, Tanaecia iapis, and the rarely sighted female of brookiana. A visit to Tapah Hills revealed more wonders, including the impressive Satyrine Neorina lowii, the golden Birdwing Triodes helena, the spectacular 5-bar Swordtail Pathysa antiphates, and several transparent Dragontails Lamproptera meges, which resembled dragonflies in flight.
Venturing up to the Cameron Highlands, we encountered the rare cloudforest Danaine Parantica sita and witnessed a gathering of Long-tailed Blues Lampides boeticus. The weather, though dull and cool, didn’t dampen our spirits as we continued our exploration. En route back to Tapah, a serendipitous encounter with an Oriental Deaths Head Hawkmoth larva added to our excitement.
Our journey then led us to the Malaysian province of Sabah, on Borneo, where vast expanses of lowland were covered in oil palm plantations. However, amidst this, we found solace in the beautiful Danum Valley, one of the last bastions of natural forest. Though the entomological findings were not as abundant as expected, the rich biodiversity of the area didn’t disappoint. Butterfly sightings included Papilio nephelus, Papilio memnon, Euploea sylvester, Hebomoia glaucippe, and many more. Beyond butterflies, our encounters with orangutans, wild boars, and monitor lizards added to the allure of the region.
Our final stop was Mulu National Park in Sarawak, a province still boasting vast areas of untouched forest. Trails to Clearwater Cave and Deer Cave treated us to sightings of various butterflies, including the lovely Papilionid Pathysa delessertii and the Plum Judy Abisara saturata. Along the plank-walk to Deer Cave, we marveled at the graceful Tree Nymph Idea lynceus, circling around us in a mesmerizing dance.
Overall, our journey through West Malaysia and Borneo in August 2004 was a testament to the region’s rich biodiversity and natural wonders, leaving us with unforgettable memories and a deep appreciation for these fascinating landscapes.
|
xFamily |
subfamily |
Genus |
species |
|
Hesperiidae |
Hesperiinae |
Aides |
duma |
|
Hesperiidae |
Hesperiinae |
Callimormus |
radiola |
|
Hesperiidae |
Hesperiinae |
Decinea |
percosius |
|
Hesperiidae |
Hesperiinae |
Dubiella |
dubius |
|
Hesperiidae |
Hesperiinae |
Panoquina |
evadnes |
|
Hesperiidae |
Hesperiinae |
Panoquina |
fusina |
|
Hesperiidae |
Hesperiinae |
Parphorus |
decora |
|
Hesperiidae |
Hesperiinae |
Parphorus |
storax |
|
Hesperiidae |
Hesperiinae |
Pellicia |
vecina |
|
Hesperiidae |
Hesperiinae |
Phanes |
aletes |
|
Hesperiidae |
Hesperiinae |
Polites |
vibex |
|
Hesperiidae |
Hesperiinae |
Pyrrhopygopsis |
romula |
|
Hesperiidae |
Hesperiinae |
Saliana |
nr. longirostris |
|
Hesperiidae |
Hesperiinae |
Tirynthia |
conflua |
|
Hesperiidae |
Hesperiinae |
Vehilius |
stictomenes |
|
Hesperiidae |
Hesperiinae |
Zenis |
jebus |
|
Hesperiidae |
Pyrginae |
Achlyodes |
busirus |
|
Hesperiidae |
Pyrginae |
Achlyodes |
mithridates |
|
Hesperiidae |
Pyrginae |
Antigonus |
liborius |
|
Hesperiidae |
Pyrginae |
Astraptes |
anaphus |
|
Hesperiidae |
Pyrginae |
Astraptes |
fulgerator |
|
Hesperiidae |
Pyrginae |
Bolla |
atahuallpai |
|
Hesperiidae |
Pyrginae |
Bolla |
cupreiceps |
|
Hesperiidae |
Pyrginae |
Bungalotis |
astylos |
|
Hesperiidae |
Pyrginae |
Celaenorrhinus |
monartus |
|
Hesperiidae |
Pyrginae |
Camptopleura |
theramenes |
|
Hesperiidae |
Pyrginae |
Carrhenes |
bamba |
|
Hesperiidae |
Pyrginae |
Cycloglypha |
thrasibulus |
|
Hesperiidae |
Pyrginae |
Cycloglypha |
tisias |
|
Hesperiidae |
Pyrginae |
Cyclosemia |
herennius |
|
Hesperiidae |
Pyrginae |
Ebrietas |
anacreon |
|
Hesperiidae |
Pyrginae |
Gorgythion |
alcandra |
|
Hesperiidae |
Pyrginae |
Grais |
stigmaticus |
|
Hesperiidae |
Pyrginae |
Helias |
phalaenoides |
|
Hesperiidae |
Pyrginae |
Heliopetes |
petrus |
|
Hesperiidae |
Pyrginae |
Hyalothyrus |
infernalis |
|
Hesperiidae |
Pyrginae |
Hyalothyrus |
leucomelas |
|
Hesperiidae |
Pyrginae |
Milanion |
leucaspis |
|
Hesperiidae |
Pyrginae |
Milanion |
hemes |
|
Hesperiidae |
Pyrginae |
Mylon |
maimon |
|
Hesperiidae |
Pyrginae |
Pellicia |
vecina |
|
Hesperiidae |
Pyrginae |
Polyctor |
tensa |
|
Hesperiidae |
Pyrginae |
Polythrix |
auginus |
|
Hesperiidae |
Pyrginae |
Polythrix |
octomaculata |
|
Hesperiidae |
Pyrginae |
Porphyrogenes |
vulpecula |
|
Hesperiidae |
Pyrginae |
Pyrgus |
oileus |
|
Hesperiidae |
Pyrginae |
Pythonides |
jovianus |
|
Hesperiidae |
Pyrginae |
Sostrata |
festiva |
|
Hesperiidae |
Pyrginae |
Telemiades |
avitus |
|
Hesperiidae |
Pyrginae |
Telemiades |
delalanda |
|
Hesperiidae |
Pyrginae |
Urbanus |
doryssus |
|
Hesperiidae |
Pyrginae |
Urbanus |
simplicius |
|
Hesperiidae |
Pyrginae |
Urbanus |
virescens |
|
Hesperiidae |
Pyrrhopyginae |
Croniades |
pieria |
|
Hesperiidae |
Pyrrhopyginae |
Jemadia |
hewitsonii |
|
Hesperiidae |
Pyrrhopyginae |
Jemadia |
hospita |
|
Hesperiidae |
Pyrrhopyginae |
Jemadia |
sosia |
|
Hesperiidae |
Pyrrhopyginae |
Parelbella |
ahira |
|
Hesperiidae |
Pyrrhopyginae |
Pyrrhopyge |
phidias |
|
Hesperiidae |
Pyrrhopyginae |
Pyrrhopyge |
proculus |
|
Lycaenidae |
Polyommatinae |
Hemiargus |
hanno |
|
Lycaenidae |
Polyommatinae |
Leptotes |
cassius |
|
Lycaenidae |
Theclinae |
Arawacus |
separata |
|
Lycaenidae |
Theclinae |
Arcas |
imperialis |
|
Lycaenidae |
Theclinae |
Calycopis |
nr caulonia |
|
Lycaenidae |
Theclinae |
Celmia |
celmus |
|
Lycaenidae |
Theclinae |
Chalybis |
hassan |
|
Lycaenidae |
Theclinae |
Electrostrymon |
mathewi |
|
Lycaenidae |
Theclinae |
Iaspis |
castitas |
|
Lycaenidae |
Theclinae |
Janthecla |
rocena |
|
Lycaenidae |
Theclinae |
Janthecla |
sista |
|
Lycaenidae |
Theclinae |
Michaelus |
ira |
|
Lycaenidae |
Theclinae |
Ministrymon |
zilda |
|
Lycaenidae |
Theclinae |
Mithras |
nautes |
|
Lycaenidae |
Theclinae |
Ocaria |
ocrisia |
|
Lycaenidae |
Theclinae |
Ocaria |
thales |
|
Lycaenidae |
Theclinae |
Olynthus |
essus |
|
Lycaenidae |
Theclinae |
Ostrinotes |
tarena |
|
Lycaenidae |
Theclinae |
Panthiades |
bitias |
|
Lycaenidae |
Theclinae |
Rekoa |
stagira |
|
Lycaenidae |
Theclinae |
Symbiopsis |
sp. nov |
|
Lycaenidae |
Theclinae |
Theclopsis |
lydus |
|
Lycaenidae |
Theclinae |
Theritas |
hemon |
|
Lycaenidae |
Theclinae |
Thestius |
pholeus |
|
Lycaenidae |
Theclinae |
Tmolus |
echion |
|
Nymphalidae |
Apaturinae |
Doxocopa |
agathina |
|
Nymphalidae |
Apaturinae |
Doxocopa |
lavinia |
|
Nymphalidae |
Biblidinae |
Callicore |
astarte |
|
Nymphalidae |
Biblidinae |
Callicore |
cynosura |
|
Nymphalidae |
Biblidinae |
Callicore |
hystaspes |
|
Nymphalidae |
Biblidinae |
Callicore |
texa |
|
Nymphalidae |
Biblidinae |
Catonephele |
acontius |
|
Nymphalidae |
Biblidinae |
Catonephele |
numilia |
|
Nymphalidae |
Biblidinae |
Diaethria |
candrena |
|
Nymphalidae |
Biblidinae |
Diaethria |
clymena |
|
Nymphalidae |
Biblidinae |
Dynamine |
agacles |
|
Nymphalidae |
Biblidinae |
Dynamine |
arene |
|
Nymphalidae |
Biblidinae |
Dynamine |
theseus |
|
Nymphalidae |
Biblidinae |
Ectima |
iona |
|
Nymphalidae |
Biblidinae |
Eunica |
pusilla |
|
Nymphalidae |
Biblidinae |
Hamadryas |
amphinome |
|
Nymphalidae |
Biblidinae |
Hamadryas |
chloe |
|
Nymphalidae |
Biblidinae |
Hamadryas |
februa |
|
Nymphalidae |
Biblidinae |
Hamadryas |
feronia |
|
Nymphalidae |
Biblidinae |
Hamadryas |
laodamia |
|
Nymphalidae |
Biblidinae |
Hamadryas |
velutina |
|
Nymphalidae |
Biblidinae |
Marpesia |
berania |
|
Nymphalidae |
Biblidinae |
Marpesia |
chiron |
|
Nymphalidae |
Biblidinae |
Marpesia |
furcula |
|
Nymphalidae |
Biblidinae |
Marpesia |
orsilochus |
|
Nymphalidae |
Biblidinae |
Marpesia |
petreus |
|
Nymphalidae |
Biblidinae |
Marpesia |
themistocles |
|
Nymphalidae |
Biblidinae |
Nessaea |
obrinus |
|
Nymphalidae |
Biblidinae |
Nica |
flavilla |
|
Nymphalidae |
Biblidinae |
Peria |
lamis |
|
Nymphalidae |
Biblidinae |
Pyrrhogyra |
amphiro |
|
Nymphalidae |
Biblidinae |
Pyrrhogyra |
crameri |
|
Nymphalidae |
Biblidinae |
Temenis |
laothoe |
|
Nymphalidae |
Biblidinae |
Temenis |
pulchra |
|
Nymphalidae |
Biblidinae |
Vila |
emilia |
|
Nymphalidae |
Charaxinae |
Agrias |
claudina |
|
Nymphalidae |
Charaxinae |
Archaeoprepona |
demophon |
|
Nymphalidae |
Charaxinae |
Archaeoprepona |
demophoon |
|
Nymphalidae |
Charaxinae |
Consul |
fabius |
|
Nymphalidae |
Charaxinae |
Fountainea |
ryphea |
|
Nymphalidae |
Charaxinae |
Memphis |
moruus |
|
Nymphalidae |
Charaxinae |
Siderone |
galanthis |
|
Nymphalidae |
Danainae |
Lycorea |
halia |
|
Nymphalidae |
Heliconiinae |
Dryas |
iulia |
|
Nymphalidae |
Heliconiinae |
Eueides |
aliphera |
|
Nymphalidae |
Heliconiinae |
Heliconius |
burneyi |
|
Nymphalidae |
Heliconiinae |
Heliconius |
erato |
|
Nymphalidae |
Heliconiinae |
Heliconius |
sara |
|
Nymphalidae |
Heliconiinae |
Heliconius |
wallacei |
|
Nymphalidae |
Heliconiinae |
Philaethria |
dido |
|
Nymphalidae |
Ithomiinae |
Aeria |
elara |
|
Nymphalidae |
Ithomiinae |
Callithomia |
lenea |
|
Nymphalidae |
Limenitidinae |
Adelpha |
basiloides |
|
Nymphalidae |
Limenitidinae |
Adelpha |
boeotia |
|
Nymphalidae |
Limenitidinae |
Adelpha |
cocala |
|
Nymphalidae |
Limenitidinae |
Adelpha |
epione |
|
Nymphalidae |
Limenitidinae |
Adelpha |
erotia |
|
Nymphalidae |
Limenitidinae |
Adelpha |
iphiclus |
|
Nymphalidae |
Limenitidinae |
Adelpha |
mesentina |
|
Nymphalidae |
Limenitidinae |
Adelpha |
messana |
|
Nymphalidae |
Limenitidinae |
Adelpha |
serpa |
|
Nymphalidae |
Limenitidinae |
Adelpha |
thesprotia |
|
Nymphalidae |
Limenitidinae |
Adelpha |
thoasa |
|
Nymphalidae |
Morphinae |
Antirrhea |
philaretes |
|
Nymphalidae |
Morphinae |
Antirrhea |
taygetina |
|
Nymphalidae |
Morphinae |
Bia |
actorion |
|
Nymphalidae |
Morphinae |
Caligo |
brasiliensis |
|
Nymphalidae |
Morphinae |
Caligo |
eurilochus |
|
Nymphalidae |
Morphinae |
Caligo |
idomeneus |
|
Nymphalidae |
Morphinae |
Caligo |
illioneus |
|
Nymphalidae |
Morphinae |
Catoblepia |
berecynthia |
|
Nymphalidae |
Morphinae |
Catoblepia |
soranus |
|
Nymphalidae |
Morphinae |
Eryphanis |
automedon |
|
Nymphalidae |
Morphinae |
Morpho |
achilles |
|
Nymphalidae |
Morphinae |
Morpho |
deidamia |
|
Nymphalidae |
Morphinae |
Morpho |
helenor |
|
Nymphalidae |
Morphinae |
Morpho |
meneleus |
|
Nymphalidae |
Morphinae |
Morpho |
rhetenor |
|
Nymphalidae |
Morphinae |
Opoptera |
aorsa |
|
Nymphalidae |
Morphinae |
Opsiphanes |
cassiae |
|
Nymphalidae |
Morphinae |
Opsiphanes |
invirae |
|
Nymphalidae |
Nymphalinae |
Anartia |
amathea |
|
Nymphalidae |
Nymphalinae |
Baeotus |
beotus |
|
Nymphalidae |
Nymphalinae |
Baeotus |
deucalion |
|
Nymphalidae |
Nymphalinae |
Baeotus |
japetus |
|
Nymphalidae |
Nymphalinae |
Colobura |
annulata |
|
Nymphalidae |
Nymphalinae |
Colobura |
dirce |
|
Nymphalidae |
Nymphalinae |
Eresia |
clio |
|
Nymphalidae |
Nymphalinae |
Historis |
odius |
|
Nymphalidae |
Nymphalinae |
Hypanartia |
lethe |
|
Nymphalidae |
Nymphalinae |
Ortilia |
gentina |
|
Nymphalidae |
Nymphalinae |
Siproeta |
stelenes |
|
Nymphalidae |
Nymphalinae |
Tegosa |
claudina |
|
Nymphalidae |
Nymphalinae |
Telenassa |
teletusa |
|
Nymphalidae |
Nymphalinae |
Tigridia |
acesta |
|
Nymphalidae |
Satyrinae |
Amphidecta |
calliomma |
|
Nymphalidae |
Satyrinae |
Caeruleuptychia |
brixius |
|
Nymphalidae |
Satyrinae |
Caeruleuptychia |
coelestis |
|
Nymphalidae |
Satyrinae |
Chloreuptychia |
agatha |
|
Nymphalidae |
Satyrinae |
Chloreuptychia |
arnaca |
|
Nymphalidae |
Satyrinae |
Chloreuptychia |
herseis |
|
Nymphalidae |
Satyrinae |
Cissia |
myncea |
|
Nymphalidae |
Satyrinae |
Cissia |
penelope |
|
Nymphalidae |
Satyrinae |
Cissia |
terrestris |
|
Nymphalidae |
Satyrinae |
Cissia |
pompilia |
|
Nymphalidae |
Satyrinae |
Euptychia |
mollina |
|
Nymphalidae |
Satyrinae |
Euptychia |
westwoodi |
|
Nymphalidae |
Satyrinae |
Haetera |
piera |
|
Nymphalidae |
Satyrinae |
Harjesia |
griseola |
|
Nymphalidae |
Satyrinae |
Hermeuptychia |
hermes |
|
Nymphalidae |
Satyrinae |
Magneuptychia |
iris |
|
Nymphalidae |
Satyrinae |
Magneuptychia |
mimas |
|
Nymphalidae |
Satyrinae |
Magneuptychia |
ocypete |
|
Nymphalidae |
Satyrinae |
Pareuptychia |
ocirrhoe |
|
Nymphalidae |
Satyrinae |
Pierella |
hyalinus |
|
Nymphalidae |
Satyrinae |
Pierella |
lamia |
|
Nymphalidae |
Satyrinae |
Pierella |
lena |
|
Nymphalidae |
Satyrinae |
Posttaygetis |
penelea |
|
Nymphalidae |
Satyrinae |
Pseudodebis |
valentina |
|
Nymphalidae |
Satyrinae |
Taygetis |
angulosa |
|
Nymphalidae |
Satyrinae |
Taygetis |
cleopatra |
|
Nymphalidae |
Satyrinae |
Taygetis |
laches |
|
Nymphalidae |
Satyrinae |
Taygetis |
mermeria |
|
Nymphalidae |
Satyrinae |
Taygetis |
rufomarginata |
|
Nymphalidae |
Satyrinae |
Taygetis |
ypthima |
|
Nymphalidae |
Satyrinae |
Taygetomorpha |
celia |
|
Papilionidae |
Papilioninae |
Battus |
crassus |
|
Papilionidae |
Papilioninae |
Battus |
lycidas |
|
Papilionidae |
Papilioninae |
Heraclides |
astyalus |
|
Papilionidae |
Papilioninae |
Heraclides |
torquatus |
|
Papilionidae |
Papilioninae |
Mimoides |
ariarathes |
|
Papilionidae |
Papilioninae |
Mimoides |
euryleon |
|
Papilionidae |
Papilioninae |
Mimoides |
pausanias |
|
Papilionidae |
Papilioninae |
Parides |
sesostris |
|
Papilionidae |
Papilioninae |
Protographium |
agesilaus |
|
Pieridae |
Coliadinae |
Anteos |
menippe |
|
Pieridae |
Coliadinae |
Aphrissa |
statira |
|
Pieridae |
Coliadinae |
Eurema |
albula |
|
Pieridae |
Coliadinae |
Eurema |
elathea |
|
Pieridae |
Coliadinae |
Leucidia |
elvina |
|
Pieridae |
Coliadinae |
Phoebis |
argante |
|
Pieridae |
Coliadinae |
Phoebis |
sennae |
|
Pieridae |
Coliadinae |
Pyrisitia |
venusta |
|
Pieridae |
Coliadinae |
Rhabdodryas |
trite |
|
Pieridae |
Dismorphiinae |
Enantia |
lina |
|
Pieridae |
Dismorphiinae |
Patia |
orise |
|
Pieridae |
Pierinae |
Glutophrissa |
drusilla |
|
Pieridae |
Pierinae |
Melete |
lycimnia |
|
Riodinidae |
Euselasiinae |
Euselasia |
alcmena |
|
Riodinidae |
Euselasiinae |
Euselasia |
angulata |
|
Riodinidae |
Euselasiinae |
Euselasia |
clesa |
|
Riodinidae |
Euselasiinae |
Euselasia |
clithra |
|
Riodinidae |
Euselasiinae |
Euselasia |
euhemerus |
|
Riodinidae |
Euselasiinae |
Euselasia |
euriteus |
|
Riodinidae |
Euselasiinae |
Euselasia |
phedica |
|
Riodinidae |
Euselasiinae |
Euselasia |
toppini |
|
Riodinidae |
Riodininae |
Alesa |
lipara |
|
Riodinidae |
Riodininae |
Amarynthis |
meneria |
|
Riodinidae |
Riodininae |
Ancyluris |
meliboeus |
|
Riodinidae |
Riodininae |
Anteros |
renaldus |
|
Riodinidae |
Riodininae |
Argyrogrammana |
sticheli |
|
Riodinidae |
Riodininae |
Aricoris |
epulus |
|
Riodinidae |
Riodininae |
Calospila |
emylius |
|
Riodinidae |
Riodininae |
Caria |
castalia |
|
Riodinidae |
Riodininae |
Caria |
mantinea |
|
Riodinidae |
Riodininae |
Caria |
trochilus |
|
Riodinidae |
Riodininae |
Chalodetta |
theodora |
|
Riodinidae |
Riodininae |
Detritivora |
cleonus |
|
Riodinidae |
Riodininae |
Detritivora |
gynaea |
|
Riodinidae |
Riodininae |
Echenais |
thelephus |
|
Riodinidae |
Riodininae |
Echydna |
punctata |
|
Riodinidae |
Riodininae |
Emesis |
eurydice |
|
Riodinidae |
Riodininae |
Emesis |
fastidiosa |
|
Riodinidae |
Riodininae |
Emesis |
lucinda |
|
Riodinidae |
Riodininae |
Emesis |
mandana |
|
Riodinidae |
Riodininae |
Emesis |
ocypore |
|
Riodinidae |
Riodininae |
Emesis |
temesa |
|
Riodinidae |
Riodininae |
Eurybia |
franciscana |
|
Riodinidae |
Riodininae |
Eurybia |
molochina |
|
Riodinidae |
Riodininae |
Eurybia |
nicaeus |
|
Riodinidae |
Riodininae |
Lasaia |
agesilas |
|
Riodinidae |
Riodininae |
Lasaia |
arsis |
|
Riodinidae |
Riodininae |
Lemonias |
zygia |
|
Riodinidae |
Riodininae |
Mesene |
epaphus |
|
Riodinidae |
Riodininae |
Mesene |
phareus |
|
Riodinidae |
Riodininae |
Mesosemia |
menoetes |
|
Riodinidae |
Riodininae |
Mesosemia |
minos |
|
Riodinidae |
Riodininae |
Mesosemia |
nyctea |
|
Riodinidae |
Riodininae |
Monethe |
albertus |
|
Riodinidae |
Riodininae |
Notheme |
erota |
|
Riodinidae |
Riodininae |
Nymphidium |
caricae |
|
Riodinidae |
Riodininae |
Nymphidium |
leucosia |
|
Riodinidae |
Riodininae |
Nymphidium |
olinda |
|
Riodinidae |
Riodininae |
Parcella |
amarynthina |
|
Riodinidae |
Riodininae |
Protonymphidia |
senta |
|
Riodinidae |
Riodininae |
Rhetus |
periander |
|
Riodinidae |
Riodininae |
Riodina |
lysippus |
|
Riodinidae |
Riodininae |
Sarota |
gyas |
|
Riodinidae |
Riodininae |
Semomesia |
capanea |
|
Riodinidae |
Riodininae |
Semomesia |
marisa |
|
Riodinidae |
Riodininae |
Stalachtis |
calliope |
|
Riodinidae |
Riodininae |
Synargis |
galena |
|
Riodinidae |
Riodininae |
Synargis |
orestessa |