Trip Reports – Learn Butterflies

Malaysia & Borneo 2004

learnbutterflies.com — Tue, 19 Mar 2024 01:54:46 +0000

In August 2004, a non-profit small group trip organized and led by Adrian Hoskins took us on an enthralling journey through the diverse landscapes of West Malaysia and Borneo.

Our adventure began in Tapah on the Malayan peninsula, where we encountered the magnificent Rajah Brook’s Birdwing Trogonoptera brookiana in Kuala Woh Park. Stunning mud-puddling groups of these creatures, with as many as 30 individuals, adorned the landscape with their iridescent green and black wings outstretched. Edging down the riverbank, I found myself amidst these captivating creatures, experiencing absolute bliss.

While most other species at Kuala Woh were found in ones and twos, we encountered some notable finds including the long-tailed, bright orange hairstreak Eooxylides tharis, the gorgeous Euploea radamanthus, Appias lyncida, Tanaecia iapis, and the rarely sighted female of brookiana. A visit to Tapah Hills revealed more wonders, including the impressive Satyrine Neorina lowii, the golden Birdwing Triodes helena, the spectacular 5-bar Swordtail Pathysa antiphates, and several transparent Dragontails Lamproptera meges, which resembled dragonflies in flight.

Venturing up to the Cameron Highlands, we encountered the rare cloudforest Danaine Parantica sita and witnessed a gathering of Long-tailed Blues Lampides boeticus. The weather, though dull and cool, didn’t dampen our spirits as we continued our exploration. En route back to Tapah, a serendipitous encounter with an Oriental Deaths Head Hawkmoth larva added to our excitement.

Our journey then led us to the Malaysian province of Sabah, on Borneo, where vast expanses of lowland were covered in oil palm plantations. However, amidst this, we found solace in the beautiful Danum Valley, one of the last bastions of natural forest. Though the entomological findings were not as abundant as expected, the rich biodiversity of the area didn’t disappoint. Butterfly sightings included Papilio nephelus, Papilio memnon, Euploea sylvester, Hebomoia glaucippe, and many more. Beyond butterflies, our encounters with orangutans, wild boars, and monitor lizards added to the allure of the region.

Our final stop was Mulu National Park in Sarawak, a province still boasting vast areas of untouched forest. Trails to Clearwater Cave and Deer Cave treated us to sightings of various butterflies, including the lovely Papilionid Pathysa delessertii and the Plum Judy Abisara saturata. Along the plank-walk to Deer Cave, we marveled at the graceful Tree Nymph Idea lynceus, circling around us in a mesmerizing dance.

Overall, our journey through West Malaysia and Borneo in August 2004 was a testament to the region’s rich biodiversity and natural wonders, leaving us with unforgettable memories and a deep appreciation for these fascinating landscapes.

xFamily	subfamily	Genus	species
Hesperiidae	Hesperiinae	Aides	duma
Hesperiidae	Hesperiinae	Callimormus	radiola
Hesperiidae	Hesperiinae	Decinea	percosius
Hesperiidae	Hesperiinae	Dubiella	dubius
Hesperiidae	Hesperiinae	Panoquina	evadnes
Hesperiidae	Hesperiinae	Panoquina	fusina
Hesperiidae	Hesperiinae	Parphorus	decora
Hesperiidae	Hesperiinae	Parphorus	storax
Hesperiidae	Hesperiinae	Pellicia	vecina
Hesperiidae	Hesperiinae	Phanes	aletes
Hesperiidae	Hesperiinae	Polites	vibex
Hesperiidae	Hesperiinae	Pyrrhopygopsis	romula
Hesperiidae	Hesperiinae	Saliana	nr. longirostris
Hesperiidae	Hesperiinae	Tirynthia	conflua
Hesperiidae	Hesperiinae	Vehilius	stictomenes
Hesperiidae	Hesperiinae	Zenis	jebus
Hesperiidae	Pyrginae	Achlyodes	busirus
Hesperiidae	Pyrginae	Achlyodes	mithridates
Hesperiidae	Pyrginae	Antigonus	liborius
Hesperiidae	Pyrginae	Astraptes	anaphus
Hesperiidae	Pyrginae	Astraptes	fulgerator
Hesperiidae	Pyrginae	Bolla	atahuallpai
Hesperiidae	Pyrginae	Bolla	cupreiceps
Hesperiidae	Pyrginae	Bungalotis	astylos
Hesperiidae	Pyrginae	Celaenorrhinus	monartus
Hesperiidae	Pyrginae	Camptopleura	theramenes
Hesperiidae	Pyrginae	Carrhenes	bamba
Hesperiidae	Pyrginae	Cycloglypha	thrasibulus
Hesperiidae	Pyrginae	Cycloglypha	tisias
Hesperiidae	Pyrginae	Cyclosemia	herennius
Hesperiidae	Pyrginae	Ebrietas	anacreon
Hesperiidae	Pyrginae	Gorgythion	alcandra
Hesperiidae	Pyrginae	Grais	stigmaticus
Hesperiidae	Pyrginae	Helias	phalaenoides
Hesperiidae	Pyrginae	Heliopetes	petrus
Hesperiidae	Pyrginae	Hyalothyrus	infernalis
Hesperiidae	Pyrginae	Hyalothyrus	leucomelas
Hesperiidae	Pyrginae	Milanion	leucaspis
Hesperiidae	Pyrginae	Milanion	hemes
Hesperiidae	Pyrginae	Mylon	maimon
Hesperiidae	Pyrginae	Pellicia	vecina
Hesperiidae	Pyrginae	Polyctor	tensa
Hesperiidae	Pyrginae	Polythrix	auginus
Hesperiidae	Pyrginae	Polythrix	octomaculata
Hesperiidae	Pyrginae	Porphyrogenes	vulpecula
Hesperiidae	Pyrginae	Pyrgus	oileus
Hesperiidae	Pyrginae	Pythonides	jovianus
Hesperiidae	Pyrginae	Sostrata	festiva
Hesperiidae	Pyrginae	Telemiades	avitus
Hesperiidae	Pyrginae	Telemiades	delalanda
Hesperiidae	Pyrginae	Urbanus	doryssus
Hesperiidae	Pyrginae	Urbanus	simplicius
Hesperiidae	Pyrginae	Urbanus	virescens
Hesperiidae	Pyrrhopyginae	Croniades	pieria
Hesperiidae	Pyrrhopyginae	Jemadia	hewitsonii
Hesperiidae	Pyrrhopyginae	Jemadia	hospita
Hesperiidae	Pyrrhopyginae	Jemadia	sosia
Hesperiidae	Pyrrhopyginae	Parelbella	ahira
Hesperiidae	Pyrrhopyginae	Pyrrhopyge	phidias
Hesperiidae	Pyrrhopyginae	Pyrrhopyge	proculus
Lycaenidae	Polyommatinae	Hemiargus	hanno
Lycaenidae	Polyommatinae	Leptotes	cassius
Lycaenidae	Theclinae	Arawacus	separata
Lycaenidae	Theclinae	Arcas	imperialis
Lycaenidae	Theclinae	Calycopis	nr caulonia
Lycaenidae	Theclinae	Celmia	celmus
Lycaenidae	Theclinae	Chalybis	hassan
Lycaenidae	Theclinae	Electrostrymon	mathewi
Lycaenidae	Theclinae	Iaspis	castitas
Lycaenidae	Theclinae	Janthecla	rocena
Lycaenidae	Theclinae	Janthecla	sista
Lycaenidae	Theclinae	Michaelus	ira
Lycaenidae	Theclinae	Ministrymon	zilda
Lycaenidae	Theclinae	Mithras	nautes
Lycaenidae	Theclinae	Ocaria	ocrisia
Lycaenidae	Theclinae	Ocaria	thales
Lycaenidae	Theclinae	Olynthus	essus
Lycaenidae	Theclinae	Ostrinotes	tarena
Lycaenidae	Theclinae	Panthiades	bitias
Lycaenidae	Theclinae	Rekoa	stagira
Lycaenidae	Theclinae	Symbiopsis	sp. nov
Lycaenidae	Theclinae	Theclopsis	lydus
Lycaenidae	Theclinae	Theritas	hemon
Lycaenidae	Theclinae	Thestius	pholeus
Lycaenidae	Theclinae	Tmolus	echion
Nymphalidae	Apaturinae	Doxocopa	agathina
Nymphalidae	Apaturinae	Doxocopa	lavinia
Nymphalidae	Biblidinae	Callicore	astarte
Nymphalidae	Biblidinae	Callicore	cynosura
Nymphalidae	Biblidinae	Callicore	hystaspes
Nymphalidae	Biblidinae	Callicore	texa
Nymphalidae	Biblidinae	Catonephele	acontius
Nymphalidae	Biblidinae	Catonephele	numilia
Nymphalidae	Biblidinae	Diaethria	candrena
Nymphalidae	Biblidinae	Diaethria	clymena
Nymphalidae	Biblidinae	Dynamine	agacles
Nymphalidae	Biblidinae	Dynamine	arene
Nymphalidae	Biblidinae	Dynamine	theseus
Nymphalidae	Biblidinae	Ectima	iona
Nymphalidae	Biblidinae	Eunica	pusilla
Nymphalidae	Biblidinae	Hamadryas	amphinome
Nymphalidae	Biblidinae	Hamadryas	chloe
Nymphalidae	Biblidinae	Hamadryas	februa
Nymphalidae	Biblidinae	Hamadryas	feronia
Nymphalidae	Biblidinae	Hamadryas	laodamia
Nymphalidae	Biblidinae	Hamadryas	velutina
Nymphalidae	Biblidinae	Marpesia	berania
Nymphalidae	Biblidinae	Marpesia	chiron
Nymphalidae	Biblidinae	Marpesia	furcula
Nymphalidae	Biblidinae	Marpesia	orsilochus
Nymphalidae	Biblidinae	Marpesia	petreus
Nymphalidae	Biblidinae	Marpesia	themistocles
Nymphalidae	Biblidinae	Nessaea	obrinus
Nymphalidae	Biblidinae	Nica	flavilla
Nymphalidae	Biblidinae	Peria	lamis
Nymphalidae	Biblidinae	Pyrrhogyra	amphiro
Nymphalidae	Biblidinae	Pyrrhogyra	crameri
Nymphalidae	Biblidinae	Temenis	laothoe
Nymphalidae	Biblidinae	Temenis	pulchra
Nymphalidae	Biblidinae	Vila	emilia
Nymphalidae	Charaxinae	Agrias	claudina
Nymphalidae	Charaxinae	Archaeoprepona	demophon
Nymphalidae	Charaxinae	Archaeoprepona	demophoon
Nymphalidae	Charaxinae	Consul	fabius
Nymphalidae	Charaxinae	Fountainea	ryphea
Nymphalidae	Charaxinae	Memphis	moruus
Nymphalidae	Charaxinae	Siderone	galanthis
Nymphalidae	Danainae	Lycorea	halia
Nymphalidae	Heliconiinae	Dryas	iulia
Nymphalidae	Heliconiinae	Eueides	aliphera
Nymphalidae	Heliconiinae	Heliconius	burneyi
Nymphalidae	Heliconiinae	Heliconius	erato
Nymphalidae	Heliconiinae	Heliconius	sara
Nymphalidae	Heliconiinae	Heliconius	wallacei
Nymphalidae	Heliconiinae	Philaethria	dido
Nymphalidae	Ithomiinae	Aeria	elara
Nymphalidae	Ithomiinae	Callithomia	lenea
Nymphalidae	Limenitidinae	Adelpha	basiloides
Nymphalidae	Limenitidinae	Adelpha	boeotia
Nymphalidae	Limenitidinae	Adelpha	cocala
Nymphalidae	Limenitidinae	Adelpha	epione
Nymphalidae	Limenitidinae	Adelpha	erotia
Nymphalidae	Limenitidinae	Adelpha	iphiclus
Nymphalidae	Limenitidinae	Adelpha	mesentina
Nymphalidae	Limenitidinae	Adelpha	messana
Nymphalidae	Limenitidinae	Adelpha	serpa
Nymphalidae	Limenitidinae	Adelpha	thesprotia
Nymphalidae	Limenitidinae	Adelpha	thoasa
Nymphalidae	Morphinae	Antirrhea	philaretes
Nymphalidae	Morphinae	Antirrhea	taygetina
Nymphalidae	Morphinae	Bia	actorion
Nymphalidae	Morphinae	Caligo	brasiliensis
Nymphalidae	Morphinae	Caligo	eurilochus
Nymphalidae	Morphinae	Caligo	idomeneus
Nymphalidae	Morphinae	Caligo	illioneus
Nymphalidae	Morphinae	Catoblepia	berecynthia
Nymphalidae	Morphinae	Catoblepia	soranus
Nymphalidae	Morphinae	Eryphanis	automedon
Nymphalidae	Morphinae	Morpho	achilles
Nymphalidae	Morphinae	Morpho	deidamia
Nymphalidae	Morphinae	Morpho	helenor
Nymphalidae	Morphinae	Morpho	meneleus
Nymphalidae	Morphinae	Morpho	rhetenor
Nymphalidae	Morphinae	Opoptera	aorsa
Nymphalidae	Morphinae	Opsiphanes	cassiae
Nymphalidae	Morphinae	Opsiphanes	invirae
Nymphalidae	Nymphalinae	Anartia	amathea
Nymphalidae	Nymphalinae	Baeotus	beotus
Nymphalidae	Nymphalinae	Baeotus	deucalion
Nymphalidae	Nymphalinae	Baeotus	japetus
Nymphalidae	Nymphalinae	Colobura	annulata
Nymphalidae	Nymphalinae	Colobura	dirce
Nymphalidae	Nymphalinae	Eresia	clio
Nymphalidae	Nymphalinae	Historis	odius
Nymphalidae	Nymphalinae	Hypanartia	lethe
Nymphalidae	Nymphalinae	Ortilia	gentina
Nymphalidae	Nymphalinae	Siproeta	stelenes
Nymphalidae	Nymphalinae	Tegosa	claudina
Nymphalidae	Nymphalinae	Telenassa	teletusa
Nymphalidae	Nymphalinae	Tigridia	acesta
Nymphalidae	Satyrinae	Amphidecta	calliomma
Nymphalidae	Satyrinae	Caeruleuptychia	brixius
Nymphalidae	Satyrinae	Caeruleuptychia	coelestis
Nymphalidae	Satyrinae	Chloreuptychia	agatha
Nymphalidae	Satyrinae	Chloreuptychia	arnaca
Nymphalidae	Satyrinae	Chloreuptychia	herseis
Nymphalidae	Satyrinae	Cissia	myncea
Nymphalidae	Satyrinae	Cissia	penelope
Nymphalidae	Satyrinae	Cissia	terrestris
Nymphalidae	Satyrinae	Cissia	pompilia
Nymphalidae	Satyrinae	Euptychia	mollina
Nymphalidae	Satyrinae	Euptychia	westwoodi
Nymphalidae	Satyrinae	Haetera	piera
Nymphalidae	Satyrinae	Harjesia	griseola
Nymphalidae	Satyrinae	Hermeuptychia	hermes
Nymphalidae	Satyrinae	Magneuptychia	iris
Nymphalidae	Satyrinae	Magneuptychia	mimas
Nymphalidae	Satyrinae	Magneuptychia	ocypete
Nymphalidae	Satyrinae	Pareuptychia	ocirrhoe
Nymphalidae	Satyrinae	Pierella	hyalinus
Nymphalidae	Satyrinae	Pierella	lamia
Nymphalidae	Satyrinae	Pierella	lena
Nymphalidae	Satyrinae	Posttaygetis	penelea
Nymphalidae	Satyrinae	Pseudodebis	valentina
Nymphalidae	Satyrinae	Taygetis	angulosa
Nymphalidae	Satyrinae	Taygetis	cleopatra
Nymphalidae	Satyrinae	Taygetis	laches
Nymphalidae	Satyrinae	Taygetis	mermeria
Nymphalidae	Satyrinae	Taygetis	rufomarginata
Nymphalidae	Satyrinae	Taygetis	ypthima
Nymphalidae	Satyrinae	Taygetomorpha	celia
Papilionidae	Papilioninae	Battus	crassus
Papilionidae	Papilioninae	Battus	lycidas
Papilionidae	Papilioninae	Heraclides	astyalus
Papilionidae	Papilioninae	Heraclides	torquatus
Papilionidae	Papilioninae	Mimoides	ariarathes
Papilionidae	Papilioninae	Mimoides	euryleon
Papilionidae	Papilioninae	Mimoides	pausanias
Papilionidae	Papilioninae	Parides	sesostris
Papilionidae	Papilioninae	Protographium	agesilaus
Pieridae	Coliadinae	Anteos	menippe
Pieridae	Coliadinae	Aphrissa	statira
Pieridae	Coliadinae	Eurema	albula
Pieridae	Coliadinae	Eurema	elathea
Pieridae	Coliadinae	Leucidia	elvina
Pieridae	Coliadinae	Phoebis	argante
Pieridae	Coliadinae	Phoebis	sennae
Pieridae	Coliadinae	Pyrisitia	venusta
Pieridae	Coliadinae	Rhabdodryas	trite
Pieridae	Dismorphiinae	Enantia	lina
Pieridae	Dismorphiinae	Patia	orise
Pieridae	Pierinae	Glutophrissa	drusilla
Pieridae	Pierinae	Melete	lycimnia
Riodinidae	Euselasiinae	Euselasia	alcmena
Riodinidae	Euselasiinae	Euselasia	angulata
Riodinidae	Euselasiinae	Euselasia	clesa
Riodinidae	Euselasiinae	Euselasia	clithra
Riodinidae	Euselasiinae	Euselasia	euhemerus
Riodinidae	Euselasiinae	Euselasia	euriteus
Riodinidae	Euselasiinae	Euselasia	phedica
Riodinidae	Euselasiinae	Euselasia	toppini
Riodinidae	Riodininae	Alesa	lipara
Riodinidae	Riodininae	Amarynthis	meneria
Riodinidae	Riodininae	Ancyluris	meliboeus
Riodinidae	Riodininae	Anteros	renaldus
Riodinidae	Riodininae	Argyrogrammana	sticheli
Riodinidae	Riodininae	Aricoris	epulus
Riodinidae	Riodininae	Calospila	emylius
Riodinidae	Riodininae	Caria	castalia
Riodinidae	Riodininae	Caria	mantinea
Riodinidae	Riodininae	Caria	trochilus
Riodinidae	Riodininae	Chalodetta	theodora
Riodinidae	Riodininae	Detritivora	cleonus
Riodinidae	Riodininae	Detritivora	gynaea
Riodinidae	Riodininae	Echenais	thelephus
Riodinidae	Riodininae	Echydna	punctata
Riodinidae	Riodininae	Emesis	eurydice
Riodinidae	Riodininae	Emesis	fastidiosa
Riodinidae	Riodininae	Emesis	lucinda
Riodinidae	Riodininae	Emesis	mandana
Riodinidae	Riodininae	Emesis	ocypore
Riodinidae	Riodininae	Emesis	temesa
Riodinidae	Riodininae	Eurybia	franciscana
Riodinidae	Riodininae	Eurybia	molochina
Riodinidae	Riodininae	Eurybia	nicaeus
Riodinidae	Riodininae	Lasaia	agesilas
Riodinidae	Riodininae	Lasaia	arsis
Riodinidae	Riodininae	Lemonias	zygia
Riodinidae	Riodininae	Mesene	epaphus
Riodinidae	Riodininae	Mesene	phareus
Riodinidae	Riodininae	Mesosemia	menoetes
Riodinidae	Riodininae	Mesosemia	minos
Riodinidae	Riodininae	Mesosemia	nyctea
Riodinidae	Riodininae	Monethe	albertus
Riodinidae	Riodininae	Notheme	erota
Riodinidae	Riodininae	Nymphidium	caricae
Riodinidae	Riodininae	Nymphidium	leucosia
Riodinidae	Riodininae	Nymphidium	olinda
Riodinidae	Riodininae	Parcella	amarynthina
Riodinidae	Riodininae	Protonymphidia	senta
Riodinidae	Riodininae	Rhetus	periander
Riodinidae	Riodininae	Riodina	lysippus
Riodinidae	Riodininae	Sarota	gyas
Riodinidae	Riodininae	Semomesia	capanea
Riodinidae	Riodininae	Semomesia	marisa
Riodinidae	Riodininae	Stalachtis	calliope
Riodinidae	Riodininae	Synargis	galena
Riodinidae	Riodininae	Synargis	orestessa

Brazil August 2006

learnbutterflies.com — Tue, 19 Mar 2024 01:55:40 +0000

In August 2006, a non-profit small group tour organized and led by Adrian Hoskins embarked on an enthralling expedition to Rio Cristalino, reaffirming its status as one of the most prolific butterfly sites in the neotropics. Limited to just four participants, our group aimed to minimize trampling at mud holes and facilitate photography on narrow trails.

Our journey commenced with a long and wearying flight from London via Dallas and Sao Paulo to Cuiaba. Upon arrival, we sought respite at a city hotel for a well-deserved night’s rest before transferring to the domestic airport for our flight to Alta Floresta. After a hearty lunch, we embarked on a short boat journey from the jetty on the Rio Teles Peres to our lodge on the Rio Cristalino.

At Rio Cristalino, our experienced guide and boatman skillfully baited tree-trunks with banana and fish, drawing forth a multitude of butterflies. The trails teemed with Satyrines, Lycaenids, and Riodinids, with notable sightings including a stunning male Hamadryas velutina basking on a tree trunk. The peccary wallow emerged as a hotspot, attracting a plethora of Nymphalids, showcasing species like the striking Siderone galanthis and the brilliant Caria mantinea.

Radiating from the lodge, trails offered excellent sightings of Satyrines, including the exquisite dead-leaf mimics Taygetis angulosa and Taygetis mermeria. Brassolines, such as Caligo Owl butterflies and Catoblepia berecynthia, added to the diversity. Sarota species buzzed around banana trees, their intricate markings captivating our attention.

Explorations by longboat unveiled new butterfly species and breathtaking encounters with kingfishers. The highlight, however, was our encounter with a gigantic Anaconda, its presence evoking both awe and trepidation.

Species recorded at Rio Cristalino in August 2006 were meticulously identified by Adrian Hoskins and Tony Hoare, drawing from extensive resources and consultations with taxonomists specializing in the relevant families. This journey was a testament to the rich biodiversity and wonders of Rio Cristalino, leaving us with unforgettable memories and a deeper appreciation for the natural world.

xFamily	subfamily	Genus	species
Hesperiidae	Hesperiinae	Aides	duma
Hesperiidae	Hesperiinae	Callimormus	radiola
Hesperiidae	Hesperiinae	Decinea	percosius
Hesperiidae	Hesperiinae	Dubiella	dubius
Hesperiidae	Hesperiinae	Panoquina	evadnes
Hesperiidae	Hesperiinae	Panoquina	fusina
Hesperiidae	Hesperiinae	Parphorus	decora
Hesperiidae	Hesperiinae	Parphorus	storax
Hesperiidae	Hesperiinae	Pellicia	vecina
Hesperiidae	Hesperiinae	Phanes	aletes
Hesperiidae	Hesperiinae	Polites	vibex
Hesperiidae	Hesperiinae	Pyrrhopygopsis	romula
Hesperiidae	Hesperiinae	Saliana	nr. longirostris
Hesperiidae	Hesperiinae	Tirynthia	conflua
Hesperiidae	Hesperiinae	Vehilius	stictomenes
Hesperiidae	Hesperiinae	Zenis	jebus
Hesperiidae	Pyrginae	Achlyodes	busirus
Hesperiidae	Pyrginae	Achlyodes	mithridates
Hesperiidae	Pyrginae	Antigonus	liborius
Hesperiidae	Pyrginae	Astraptes	anaphus
Hesperiidae	Pyrginae	Astraptes	fulgerator
Hesperiidae	Pyrginae	Bolla	atahuallpai
Hesperiidae	Pyrginae	Bolla	cupreiceps
Hesperiidae	Pyrginae	Bungalotis	astylos
Hesperiidae	Pyrginae	Celaenorrhinus	monartus
Hesperiidae	Pyrginae	Camptopleura	theramenes
Hesperiidae	Pyrginae	Carrhenes	bamba
Hesperiidae	Pyrginae	Cycloglypha	thrasibulus
Hesperiidae	Pyrginae	Cycloglypha	tisias
Hesperiidae	Pyrginae	Cyclosemia	herennius
Hesperiidae	Pyrginae	Ebrietas	anacreon
Hesperiidae	Pyrginae	Gorgythion	alcandra
Hesperiidae	Pyrginae	Grais	stigmaticus
Hesperiidae	Pyrginae	Helias	phalaenoides
Hesperiidae	Pyrginae	Heliopetes	petrus
Hesperiidae	Pyrginae	Hyalothyrus	infernalis
Hesperiidae	Pyrginae	Hyalothyrus	leucomelas
Hesperiidae	Pyrginae	Milanion	leucaspis
Hesperiidae	Pyrginae	Milanion	hemes
Hesperiidae	Pyrginae	Mylon	maimon
Hesperiidae	Pyrginae	Pellicia	vecina
Hesperiidae	Pyrginae	Polyctor	tensa
Hesperiidae	Pyrginae	Polythrix	auginus
Hesperiidae	Pyrginae	Polythrix	octomaculata
Hesperiidae	Pyrginae	Porphyrogenes	vulpecula
Hesperiidae	Pyrginae	Pyrgus	oileus
Hesperiidae	Pyrginae	Pythonides	jovianus
Hesperiidae	Pyrginae	Sostrata	festiva
Hesperiidae	Pyrginae	Telemiades	avitus
Hesperiidae	Pyrginae	Telemiades	delalanda
Hesperiidae	Pyrginae	Urbanus	doryssus
Hesperiidae	Pyrginae	Urbanus	simplicius
Hesperiidae	Pyrginae	Urbanus	virescens
Hesperiidae	Pyrrhopyginae	Croniades	pieria
Hesperiidae	Pyrrhopyginae	Jemadia	hewitsonii
Hesperiidae	Pyrrhopyginae	Jemadia	hospita
Hesperiidae	Pyrrhopyginae	Jemadia	sosia
Hesperiidae	Pyrrhopyginae	Parelbella	ahira
Hesperiidae	Pyrrhopyginae	Pyrrhopyge	phidias
Hesperiidae	Pyrrhopyginae	Pyrrhopyge	proculus
Lycaenidae	Polyommatinae	Hemiargus	hanno
Lycaenidae	Polyommatinae	Leptotes	cassius
Lycaenidae	Theclinae	Arawacus	separata
Lycaenidae	Theclinae	Arcas	imperialis
Lycaenidae	Theclinae	Calycopis	nr caulonia
Lycaenidae	Theclinae	Celmia	celmus
Lycaenidae	Theclinae	Chalybis	hassan
Lycaenidae	Theclinae	Electrostrymon	mathewi
Lycaenidae	Theclinae	Iaspis	castitas
Lycaenidae	Theclinae	Janthecla	rocena
Lycaenidae	Theclinae	Janthecla	sista
Lycaenidae	Theclinae	Michaelus	ira
Lycaenidae	Theclinae	Ministrymon	zilda
Lycaenidae	Theclinae	Mithras	nautes
Lycaenidae	Theclinae	Ocaria	ocrisia
Lycaenidae	Theclinae	Ocaria	thales
Lycaenidae	Theclinae	Olynthus	essus
Lycaenidae	Theclinae	Ostrinotes	tarena
Lycaenidae	Theclinae	Panthiades	bitias
Lycaenidae	Theclinae	Rekoa	stagira
Lycaenidae	Theclinae	Symbiopsis	sp. nov
Lycaenidae	Theclinae	Theclopsis	lydus
Lycaenidae	Theclinae	Theritas	hemon
Lycaenidae	Theclinae	Thestius	pholeus
Lycaenidae	Theclinae	Tmolus	echion
Nymphalidae	Apaturinae	Doxocopa	agathina
Nymphalidae	Apaturinae	Doxocopa	lavinia
Nymphalidae	Biblidinae	Callicore	astarte
Nymphalidae	Biblidinae	Callicore	cynosura
Nymphalidae	Biblidinae	Callicore	hystaspes
Nymphalidae	Biblidinae	Callicore	texa
Nymphalidae	Biblidinae	Catonephele	acontius
Nymphalidae	Biblidinae	Catonephele	numilia
Nymphalidae	Biblidinae	Diaethria	candrena
Nymphalidae	Biblidinae	Diaethria	clymena
Nymphalidae	Biblidinae	Dynamine	agacles
Nymphalidae	Biblidinae	Dynamine	arene
Nymphalidae	Biblidinae	Dynamine	theseus
Nymphalidae	Biblidinae	Ectima	iona
Nymphalidae	Biblidinae	Eunica	pusilla
Nymphalidae	Biblidinae	Hamadryas	amphinome
Nymphalidae	Biblidinae	Hamadryas	chloe
Nymphalidae	Biblidinae	Hamadryas	februa
Nymphalidae	Biblidinae	Hamadryas	feronia
Nymphalidae	Biblidinae	Hamadryas	laodamia
Nymphalidae	Biblidinae	Hamadryas	velutina
Nymphalidae	Biblidinae	Marpesia	berania
Nymphalidae	Biblidinae	Marpesia	chiron
Nymphalidae	Biblidinae	Marpesia	furcula
Nymphalidae	Biblidinae	Marpesia	orsilochus
Nymphalidae	Biblidinae	Marpesia	petreus
Nymphalidae	Biblidinae	Marpesia	themistocles
Nymphalidae	Biblidinae	Nessaea	obrinus
Nymphalidae	Biblidinae	Nica	flavilla
Nymphalidae	Biblidinae	Peria	lamis
Nymphalidae	Biblidinae	Pyrrhogyra	amphiro
Nymphalidae	Biblidinae	Pyrrhogyra	crameri
Nymphalidae	Biblidinae	Temenis	laothoe
Nymphalidae	Biblidinae	Temenis	pulchra
Nymphalidae	Biblidinae	Vila	emilia
Nymphalidae	Charaxinae	Agrias	claudina
Nymphalidae	Charaxinae	Archaeoprepona	demophon
Nymphalidae	Charaxinae	Archaeoprepona	demophoon
Nymphalidae	Charaxinae	Consul	fabius
Nymphalidae	Charaxinae	Fountainea	ryphea
Nymphalidae	Charaxinae	Memphis	moruus
Nymphalidae	Charaxinae	Siderone	galanthis
Nymphalidae	Danainae	Lycorea	halia
Nymphalidae	Heliconiinae	Dryas	iulia
Nymphalidae	Heliconiinae	Eueides	aliphera
Nymphalidae	Heliconiinae	Heliconius	burneyi
Nymphalidae	Heliconiinae	Heliconius	erato
Nymphalidae	Heliconiinae	Heliconius	sara
Nymphalidae	Heliconiinae	Heliconius	wallacei
Nymphalidae	Heliconiinae	Philaethria	dido
Nymphalidae	Ithomiinae	Aeria	elara
Nymphalidae	Ithomiinae	Callithomia	lenea
Nymphalidae	Limenitidinae	Adelpha	basiloides
Nymphalidae	Limenitidinae	Adelpha	boeotia
Nymphalidae	Limenitidinae	Adelpha	cocala
Nymphalidae	Limenitidinae	Adelpha	epione
Nymphalidae	Limenitidinae	Adelpha	erotia
Nymphalidae	Limenitidinae	Adelpha	iphiclus
Nymphalidae	Limenitidinae	Adelpha	mesentina
Nymphalidae	Limenitidinae	Adelpha	messana
Nymphalidae	Limenitidinae	Adelpha	serpa
Nymphalidae	Limenitidinae	Adelpha	thesprotia
Nymphalidae	Limenitidinae	Adelpha	thoasa
Nymphalidae	Morphinae	Antirrhea	philaretes
Nymphalidae	Morphinae	Antirrhea	taygetina
Nymphalidae	Morphinae	Bia	actorion
Nymphalidae	Morphinae	Caligo	brasiliensis
Nymphalidae	Morphinae	Caligo	eurilochus
Nymphalidae	Morphinae	Caligo	idomeneus
Nymphalidae	Morphinae	Caligo	illioneus
Nymphalidae	Morphinae	Catoblepia	berecynthia
Nymphalidae	Morphinae	Catoblepia	soranus
Nymphalidae	Morphinae	Eryphanis	automedon
Nymphalidae	Morphinae	Morpho	achilles
Nymphalidae	Morphinae	Morpho	deidamia
Nymphalidae	Morphinae	Morpho	helenor
Nymphalidae	Morphinae	Morpho	meneleus
Nymphalidae	Morphinae	Morpho	rhetenor
Nymphalidae	Morphinae	Opoptera	aorsa
Nymphalidae	Morphinae	Opsiphanes	cassiae
Nymphalidae	Morphinae	Opsiphanes	invirae
Nymphalidae	Nymphalinae	Anartia	amathea
Nymphalidae	Nymphalinae	Baeotus	beotus
Nymphalidae	Nymphalinae	Baeotus	deucalion
Nymphalidae	Nymphalinae	Baeotus	japetus
Nymphalidae	Nymphalinae	Colobura	annulata
Nymphalidae	Nymphalinae	Colobura	dirce
Nymphalidae	Nymphalinae	Eresia	clio
Nymphalidae	Nymphalinae	Historis	odius
Nymphalidae	Nymphalinae	Hypanartia	lethe
Nymphalidae	Nymphalinae	Ortilia	gentina
Nymphalidae	Nymphalinae	Siproeta	stelenes
Nymphalidae	Nymphalinae	Tegosa	claudina
Nymphalidae	Nymphalinae	Telenassa	teletusa
Nymphalidae	Nymphalinae	Tigridia	acesta
Nymphalidae	Satyrinae	Amphidecta	calliomma
Nymphalidae	Satyrinae	Caeruleuptychia	brixius
Nymphalidae	Satyrinae	Caeruleuptychia	coelestis
Nymphalidae	Satyrinae	Chloreuptychia	agatha
Nymphalidae	Satyrinae	Chloreuptychia	arnaca
Nymphalidae	Satyrinae	Chloreuptychia	herseis
Nymphalidae	Satyrinae	Cissia	myncea
Nymphalidae	Satyrinae	Cissia	penelope
Nymphalidae	Satyrinae	Cissia	terrestris
Nymphalidae	Satyrinae	Cissia	pompilia
Nymphalidae	Satyrinae	Euptychia	mollina
Nymphalidae	Satyrinae	Euptychia	westwoodi
Nymphalidae	Satyrinae	Haetera	piera
Nymphalidae	Satyrinae	Harjesia	griseola
Nymphalidae	Satyrinae	Hermeuptychia	hermes
Nymphalidae	Satyrinae	Magneuptychia	iris
Nymphalidae	Satyrinae	Magneuptychia	mimas
Nymphalidae	Satyrinae	Magneuptychia	ocypete
Nymphalidae	Satyrinae	Pareuptychia	ocirrhoe
Nymphalidae	Satyrinae	Pierella	hyalinus
Nymphalidae	Satyrinae	Pierella	lamia
Nymphalidae	Satyrinae	Pierella	lena
Nymphalidae	Satyrinae	Posttaygetis	penelea
Nymphalidae	Satyrinae	Pseudodebis	valentina
Nymphalidae	Satyrinae	Taygetis	angulosa
Nymphalidae	Satyrinae	Taygetis	cleopatra
Nymphalidae	Satyrinae	Taygetis	laches
Nymphalidae	Satyrinae	Taygetis	mermeria
Nymphalidae	Satyrinae	Taygetis	rufomarginata
Nymphalidae	Satyrinae	Taygetis	ypthima
Nymphalidae	Satyrinae	Taygetomorpha	celia
Papilionidae	Papilioninae	Battus	crassus
Papilionidae	Papilioninae	Battus	lycidas
Papilionidae	Papilioninae	Heraclides	astyalus
Papilionidae	Papilioninae	Heraclides	torquatus
Papilionidae	Papilioninae	Mimoides	ariarathes
Papilionidae	Papilioninae	Mimoides	euryleon
Papilionidae	Papilioninae	Mimoides	pausanias
Papilionidae	Papilioninae	Parides	sesostris
Papilionidae	Papilioninae	Protographium	agesilaus
Pieridae	Coliadinae	Anteos	menippe
Pieridae	Coliadinae	Aphrissa	statira
Pieridae	Coliadinae	Eurema	albula
Pieridae	Coliadinae	Eurema	elathea
Pieridae	Coliadinae	Leucidia	elvina
Pieridae	Coliadinae	Phoebis	argante
Pieridae	Coliadinae	Phoebis	sennae
Pieridae	Coliadinae	Pyrisitia	venusta
Pieridae	Coliadinae	Rhabdodryas	trite
Pieridae	Dismorphiinae	Enantia	lina
Pieridae	Dismorphiinae	Patia	orise
Pieridae	Pierinae	Glutophrissa	drusilla
Pieridae	Pierinae	Melete	lycimnia
Riodinidae	Euselasiinae	Euselasia	alcmena
Riodinidae	Euselasiinae	Euselasia	angulata
Riodinidae	Euselasiinae	Euselasia	clesa
Riodinidae	Euselasiinae	Euselasia	clithra
Riodinidae	Euselasiinae	Euselasia	euhemerus
Riodinidae	Euselasiinae	Euselasia	euriteus
Riodinidae	Euselasiinae	Euselasia	phedica
Riodinidae	Euselasiinae	Euselasia	toppini
Riodinidae	Riodininae	Alesa	lipara
Riodinidae	Riodininae	Amarynthis	meneria
Riodinidae	Riodininae	Ancyluris	meliboeus
Riodinidae	Riodininae	Anteros	renaldus
Riodinidae	Riodininae	Argyrogrammana	sticheli
Riodinidae	Riodininae	Aricoris	epulus
Riodinidae	Riodininae	Calospila	emylius
Riodinidae	Riodininae	Caria	castalia
Riodinidae	Riodininae	Caria	mantinea
Riodinidae	Riodininae	Caria	trochilus
Riodinidae	Riodininae	Chalodetta	theodora
Riodinidae	Riodininae	Detritivora	cleonus
Riodinidae	Riodininae	Detritivora	gynaea
Riodinidae	Riodininae	Echenais	thelephus
Riodinidae	Riodininae	Echydna	punctata
Riodinidae	Riodininae	Emesis	eurydice
Riodinidae	Riodininae	Emesis	fastidiosa
Riodinidae	Riodininae	Emesis	lucinda
Riodinidae	Riodininae	Emesis	mandana
Riodinidae	Riodininae	Emesis	ocypore
Riodinidae	Riodininae	Emesis	temesa
Riodinidae	Riodininae	Eurybia	franciscana
Riodinidae	Riodininae	Eurybia	molochina
Riodinidae	Riodininae	Eurybia	nicaeus
Riodinidae	Riodininae	Lasaia	agesilas
Riodinidae	Riodininae	Lasaia	arsis
Riodinidae	Riodininae	Lemonias	zygia
Riodinidae	Riodininae	Mesene	epaphus
Riodinidae	Riodininae	Mesene	phareus
Riodinidae	Riodininae	Mesosemia	menoetes
Riodinidae	Riodininae	Mesosemia	minos
Riodinidae	Riodininae	Mesosemia	nyctea
Riodinidae	Riodininae	Monethe	albertus
Riodinidae	Riodininae	Notheme	erota
Riodinidae	Riodininae	Nymphidium	caricae
Riodinidae	Riodininae	Nymphidium	leucosia
Riodinidae	Riodininae	Nymphidium	olinda
Riodinidae	Riodininae	Parcella	amarynthina
Riodinidae	Riodininae	Protonymphidia	senta
Riodinidae	Riodininae	Rhetus	periander
Riodinidae	Riodininae	Riodina	lysippus
Riodinidae	Riodininae	Sarota	gyas
Riodinidae	Riodininae	Semomesia	capanea
Riodinidae	Riodininae	Semomesia	marisa
Riodinidae	Riodininae	Stalachtis	calliope
Riodinidae	Riodininae	Synargis	galena
Riodinidae	Riodininae	Synargis	orestessa

Colombia 2013

learnbutterflies.com — Tue, 19 Mar 2024 01:56:59 +0000

a non-profit small group tour organised and led by Adrian Hoskins

Trip report

The view from our lodge at Tatama national park – Adrian Hoskins

Things don’t always go according to plan in South America, but it all adds to the sense of adventure! This was our first trip to Colombia, and when we arrived we found the country was in the middle of a national strike which caused disruption and road closures on major routes. Fortunately with the aid of our excellent local guides Dora and Hernan, and our safe reliable driver Mauricio we were able to rearrange the schedule to avoid the road blocks, and a fantastic time was had by everyone!

Delicious food, comfortable accommodation, stunning scenery, gorgeous butterflies, great company and wonderful Colombian hospitality made this a very special tour!


Heliconius charithonia	Marpesia corinna	Epiphile eriopis

Heliconius cydno	Oxeoschistus simplex	Fountainea nobilis

Ithomia hyala	Adelpha serpa	Doxocopa laurentia

After spending a night at Medellin we flew to Pereira to avoid the road blocks, and then had a short drive to our first butterfly site Otun Quimbaya. Our lodge there was located amidst cloudforest at an elevation of 1900m, and most of our butterfly-watching and photography took place on the little-used forest road that stretches from the reserve entrance at 1800m up to about 2300m where the road ends. We also explored the riverbanks, and walked along the narrow trails that led from the lodge into the surrounding forest, where we saw the clearwing Satyr Pseudohaetera hypaesia, and a good selection of Ithomiine glasswings.

The latter are fascinating to watch, fluttering secretively with deep slow wing beats, and occasionally settling on leaves to pose for photographs. Downhill from the lodge we explored a grassy track at the edge of the river, which produced the stunning zebra hairstreak Arawacus leucogyna, several very attractive red and metallic blue-black Altinote species, and a huge red skipper Yanguna cosyra.

Cloudforest at Otun Quimbaya – Adrian Hoskins

Following two full days of immersive butterfly photography at Otun, we embarked on a lengthy journey to our next destination: Tatama National Park. Nestled at an elevation of 1350 meters, our lodge was situated on the fringes of this enchanting and picturesque cloud forest. Our explorations primarily unfolded along the rugged stony road, meandering upwards from the lodge to nearly 3000 meters.

Tatama lies within the Choco ecological zone, renowned for receiving some of the highest rainfall in South America. Yet, it also stands as one of the continent’s most biodiverse butterfly havens. Although intermittent cloud cover and light drizzles deterred us from venturing to the highest elevations, we found ample opportunities for discovery between 1350-1800 meters, where conditions remained relatively warm and dry.

Some among us opted for a leisurely uphill ride in vehicles, followed by a slow descent on foot back to the lodge. Others chose to trek all the way to 1800 meters and back—an exhausting but immensely rewarding endeavor. Along the way, we encountered a multitude of butterfly species, including Callicore, Diaethria, Memphis, Heliconius, Dismorphia, Catasticta, Perisama, Adelpha, Doxocopa, Mesosemia, Catonephele, Epiphile, Hypanartia, Marpesia, and Corades. Among these sightings were the majestic Owl butterfly Caligo oedipus and three resplendent blue giant Morpho species.

The highlights of our journey were undoubtedly the breathtaking pink Fountainea nobilis, the charming Necyria bellona zaneta—a diminutive metalmark adorned with metallic blue bands and crimson spots against a jet-black backdrop—and the magnificent Epiphile eriopis, boasting striking black, red, and white hues as it basked gracefully with outstretched wings on the forest track.

Cecropia tree at dawn, Tatama national park – Adrian Hoskins

Although our tour was primarily focused on butterflies, we often found ourselves distracted by the many other wonders of Tatama – stunning scenery, hummingbirds galore, beautiful orchids, moths of every shape and pattern imaginable, gorgeous dragonflies, and magnificent bugs!

Unidentified bug, Tatama national park – Adrian Hoskins

After reveling in six delightful days at Tatama, our journey led us to Rio Claro, where we spent three days at a nature resort nestled along the riverbanks at an elevation of 350 meters. While butterfly abundance in this locale was relatively modest, the diversity of species proved to be abundant. We encountered and captured photographs of numerous species not found in the higher elevations we had previously visited. Among these treasures were several exquisite metalmarks, including pristine specimens of Sarota chrysus, Ancyluris jurgensenii, Caria mantinea, Caria rhacotis, Lasaia agesilas, and Rhetus periander.

Our next stop took us to the Arrierito Piha Reserve in Antioquoia. Nestled along a tranquil forest road at an elevation of 1700 meters, the cozy lodge provided a wonderful retreat, particularly enchanting for hummingbird enthusiasts. At times, up to 40 of these mesmerizing creatures graced a favored small tree in the lodge garden.

Along the roadside, we encountered a plethora of fabulous butterflies, including the dead-leaf mimic Consul panariste, the stunning metallic blue-banded Emperors Doxocopa cyane and Doxoxopa laurentia cherubina, the charming white-banded Oressinoma typhla, and the blue-tailed metalmark Rhetus dysonii.

Traversing the trails through the cloudforest interior yielded further delights, such as the imposing and remarkably attractive Ithomiine Elzunia humboldt and the gracefully elegant longwing Heliconius clysonymus. Another noteworthy discovery was the white-banded ringlet Splendeuptychia ackeryi, a species that made its scientific debut in 2009, adding a special touch of novelty to our expedition.


Oressinoma typhla	Hypanartia lethe	Anartia jatrophae

Cyanophrys longula	Panthiades bathildis	Callicore tolima

Rhetus dysonii	Sarota chrysus	Necyria bellona

Following our two-day sojourn at the Piha reserve, we embarked on a short drive to Santa Fe de Antioquoia, where a lengthy roadside stop rewarded us with sightings of the vividly colored longwing Dryadula phaetusa, various species of Urbanus long-tailed skippers, and a multitude of attractive hairstreak species. Another nearby site yielded sightings of Monarchs Danaus plexippus and the magnificent banded hairstreak Panthiades bathildis.

Our final day was spent in the company of two friends, Juan and Martin, residents of Medellin who are deeply knowledgeable and passionate about butterfly photography. They graciously guided us to a seemingly unremarkable patch of relict cloudforest on the outskirts of Medellin, situated at an elevation of 1400 meters. To our astonishment, this humble locale proved to be one of the most fruitful sites of our entire journey! A liberal application of liquidized fish bait onto foliage swiftly attracted swarms of butterflies, including the metallic blue Heliconius erato chestertonii and several alluring Adelpha species. Yet, the butterflies that captivated us most were the distinctive Eyemark Skipper Cyclosemia leppa and the stunningly attractive furry-legged Jewelmark Anteros kupris.

In the afternoon, we ventured to another nearby site at an elevation of 1800 meters, where we were treated to sightings of the pretty green hairstreak Cyanophrys longula, numerous glasswing and tiger-patterned Ithomiines, and a variety of pronophiline Satyrs, among which was a Forsterinaria species previously unknown to science.

Pueblito de San Jose, near Medellin – Adrian Hoskins

SPECIES LIST

Species recorded in Colombia in August 2013 – list updated 28 Dec 2013

In total we saw about 430 species on the tour, of which 421 were photographed by the participants Adrian Hoskins, Peter Chadd, Peter Bygate, Les Catchick and Chris Orpin.

Most of us photographed between 200-300 species. The list which follows was compiled by Adrian Hoskins after meticulously checking all the images taken by the participants. Resources consulted included the 7 indispensable volumes of d’Abrera’s ‘Butterflies of the Neotropical region’, Keith Willmott’s ‘The Genus Adelpha’, and the Butterflies of America (Neotropical) website.

I’d like to express my thanks also to Andrew Neild, Tomasz Pyrcz, Jean Francois le Crom, Luis Miguel Constantino and Blanca Huertas for their valued assistance in determining some of the more difficult species.

Hundreds of photographs of the species seen on the tour can now be seen in the Amazon & Andes section, where you will also find detailed articles about the biology of each species.

Hesperiidae	Achylodes	busirus	heros
Hesperiidae	Achylodes	pallida
Hesperiidae	Anastrus	meliboea	meliboea
Hesperiidae	Anisochoria	pedaliodina	polysticta
Hesperiidae	Anthoptus	epictetus	epictetus
Hesperiidae	Antigonus	nearchus	nearchus
Hesperiidae	Apaustus	gracilis	gracilis
Hesperiidae	Aroma	aroma
Hesperiidae	Astraptes	anaphus
Hesperiidae	Astraptes	chiriquensis
Hesperiidae	Astraptes	fulgerator
Hesperiidae	Autochton	longipennis
Hesperiidae	Autochton	neis
Hesperiidae	Autochton	zarex
Hesperiidae	Bolla	dorsolaciniae
Hesperiidae	Bolla	giselus
Hesperiidae	Camptopleura	auxo	giselus
Hesperiidae	Camptopleura	impressus
Hesperiidae	Camptopleura	theramenes
Hesperiidae	Carrhenes	bamba	bamba
Hesperiidae	Carrhenes	canescens	canescens
Hesperiidae	Celaenorrhinus	eligius
Hesperiidae	Celaenorrhinus	suthina
Hesperiidae	Chalypyge	zereda	rufipectus
Hesperiidae	Cogia	calchas
Hesperiidae	Corticea	mendica
Hesperiidae	Cycloglypha	enega
Hesperiidae	Cycloglypha	thrasibulus	thrasibulus
Hesperiidae	Cycloglypha	tisias	tisias
Hesperiidae	Cyclosemia	leppa
Hesperiidae	Cymaenes	trebius
Hesperiidae	Cymaenes	tripunctus	theogenis
Hesperiidae	Dalla	calima
Hesperiidae	Dalla	eryonas
Hesperiidae	Dalla	frater
Hesperiidae	Dalla	plancus	plancus
Hesperiidae	Enosis	achelous
Hesperiidae	Epargyreus	aspina
Hesperiidae	Falga	jeconia
Hesperiidae	Gindanes	brebisson	panaetius
Hesperiidae	Gorgythion	begga	begga
Hesperiidae	Helias	cama
Hesperiidae	Helias	godmani
Hesperiidae	Helias	phalaenoides
Hesperiidae	Heliopetes	alana
Hesperiidae	Heliopetes	arsalte	arsalte
Hesperiidae	Heliopetes	laviana	laviana
Hesperiidae	Hylephila	phyleus	phyleus
Hesperiidae	Lento	ludo
Hesperiidae	Mictris	crispus	caerula
Hesperiidae	Milanion	hemes
Hesperiidae	Mylon	lassia
Hesperiidae	Mylon	zephus
Hesperiidae	Nisoniades	hecale
Hesperiidae	Noctuana	haematospila
Hesperiidae	Noctuana	noctua
Hesperiidae	Nyctelius	nyctelius	nyctelius
Hesperiidae	Ouleus	fridericus	fridericus
Hesperiidae	Panoquina	evadnes	evadnes
Hesperiidae	Pellicia	santana
Hesperiidae	Penicula	cristina
Hesperiidae	Phocides	pigmalion	pigmalion
Hesperiidae	Polythrix	hirtius
Hesperiidae	Pompeius	pompeius
Hesperiidae	Potamanaxas	andaemon
Hesperiidae	Potamanaxas	laoma	trigga
Hesperiidae	Potamanaxas	latrea	caliadne
Hesperiidae	Potamanaxas	melicertes
Hesperiidae	Potamanaxas	quira
Hesperiidae	Pyrgus	adepta
Hesperiidae	Pyrgus	orcus
Hesperiidae	Pyrrhopyge	evansi	evansi
Hesperiidae	Pyrrhopyge	papius	papius
Hesperiidae	Pythonides	herennius	herennius
Hesperiidae	Quadrus	lugubris	ophia
Hesperiidae	Remella	remus
Hesperiidae	Saliana	hewitsoni
Hesperiidae	Saliana	salius	salius
Hesperiidae	Saliana	severus
Hesperiidae	Spioniades	abbreviata	anta
Hesperiidae	Timochreon	satyrus	satyrus
Hesperiidae	Urbanus	albimargo	albimargo
Hesperiidae	Urbanus	esta
Hesperiidae	Urbanus	proteus
Hesperiidae	Urbanus	simplicius	simplicius
Hesperiidae	Urbanus	teleus	teleus
Hesperiidae	Vettius	coryna	catargyra
Hesperiidae	Vettius	marcus
Hesperiidae	Xenophanes	tryxus	tryxus
Hesperiidae	Yanguna	cosyra
Hesperiidae	Zenis	jebus	beckeri
Hesperiidae	Zera	hosta
Lycaenidae	Arawacus	aetolus	aetolus
Lycaenidae	Arawacus	leucogyna
Lycaenidae	Arawacus	lincoides
Lycaenidae	Brangas	felderi
Lycaenidae	Calycopis	calus
Lycaenidae	Calycopis	cerata
Lycaenidae	Calycopis	demonassa
Lycaenidae	Contrafacia	imma
Lycaenidae	Cupido	comyntas	texana
Lycaenidae	Cyanophrys	longula
Lycaenidae	Erora	subflorens
Lycaenidae	Gargina	emessa
Lycaenidae	Hemiargus	hanno
Lycaenidae	Ipidecla	schausi
Lycaenidae	Laothus	gibberosa
Lycaenidae	Leptotes	bathyllos
Lycaenidae	Micandra	platyptera
Lycaenidae	Ministrymon	azia	azia
Lycaenidae	Ministrymon	una
Lycaenidae	Mithras	colombiensis
Lycaenidae	Nicolaea	heraldica
Lycaenidae	Nicolaea	pyxis
Lycaenidae	Ocaria	aholiba
Lycaenidae	Ostrinotes	keila
Lycaenidae	Panthiades	bathildis
Lycaenidae	Pseudolycaena	marsyas
Lycaenidae	Rekoa	meton
Lycaenidae	Strymon	mulucha
Lycaenidae	Strymon	serapio
Lycaenidae	Symbiopsis	tanais
Lycaenidae	Theclopsis	demea
Lycaenidae	Thereus	orasus
Lycaenidae	Theritas	augustinula
Lycaenidae	Theritas	hemon
Lycaenidae	Theritas	mavors
Lycaenidae	Tmolus	echion	echiolus
Lycaenidae	Zizula	cyna
Nymphalidae	Actinote	anteas	anteas
Nymphalidae	Actinote	pellenea	limbata
Nymphalidae	Adelpha	alala	negra
Nymphalidae	Adelpha	barnesia	leucas
Nymphalidae	Adelpha	boeotia	boeotia
Nymphalidae	Adelpha	cytherea	daguana
Nymphalidae	Adelpha	ethelda	eponina
Nymphalidae	Adelpha	justina	justina
Nymphalidae	Adelpha	lamasi
Nymphalidae	Adelpha	leucopthalma	leucopthalma
Nymphalidae	Adelpha	levona
Nymphalidae	Adelpha	lycorias	wallisii
Nymphalidae	Adelpha	melona	deborah
Nymphalidae	Adelpha	rothschildi
Nymphalidae	Adelpha	serpa	celerio
Nymphalidae	Aeria	eurimedia	latistriga
Nymphalidae	Altinote	neleus	neleus
Nymphalidae	Altinote	ozomene
Nymphalidae	Altinote	stratonice	stratonice
Nymphalidae	Amphidecta	calliomma
Nymphalidae	Anartia	amathea	amathea
Nymphalidae	Anartia	jatrophae	jatrophae
Nymphalidae	Anthanassa	acesas	castianira
Nymphalidae	Anthanassa	drusilla	drusilla
Nymphalidae	Archaeoprepona	demophon	muson
Nymphalidae	Archaeoprepona	meander	megabates
Nymphalidae	Caligo	illioneus	illioneus
Nymphalidae	Caligo	oedipus	oedipus
Nymphalidae	Callicore	atacama	manova
Nymphalidae	Callicore	lyca
Nymphalidae	Callicore	pitheas
Nymphalidae	Callicore	tolima	denina
Nymphalidae	Castilia	castilla	castilla
Nymphalidae	Castilia	eranites
Nymphalidae	Catonephele	chromis	godmani
Nymphalidae	Catonephele	numilia	esite
Nymphalidae	Catonephele	nyctimus
Nymphalidae	Ceratinia	tutia	transversa
Nymphalidae	Chlosyne	narva	bonpland
Nymphalidae	Chlosyne	theona	hondana
Nymphalidae	Cissia	labe
Nymphalidae	Colobura	annulata	annulata
Nymphalidae	Colobura	dirce	dirce
Nymphalidae	Consul	fabius
Nymphalidae	Consul	panariste	ludmilla
Nymphalidae	Corades	enyo	almo
Nymphalidae	Corades	pannonia	ploas
Nymphalidae	Dagon	pusilla
Nymphalidae	Danaus	eresimus	montezuma
Nymphalidae	Danaus	plexippus
Nymphalidae	Diaethria	clymena	marchalii
Nymphalidae	Diaethria	euclides	phlogea
Nymphalidae	Dione	juno	andicola
Nymphalidae	Dione	moneta	butleri
Nymphalidae	Dircenna	dero
Nymphalidae	Doxocopa	cyane	mexicana
Nymphalidae	Doxocopa	laurentia	cherubina
Nymphalidae	Doxocopa	pavon
Nymphalidae	Doxocopa	zunilda	felderi
Nymphalidae	Dryadula	phaetusa
Nymphalidae	Dryas	iulia	iulia
Nymphalidae	Dynamine	chryseis
Nymphalidae	Dynamine	paulina	thalassina
Nymphalidae	Dynamine	postverta	postverta
Nymphalidae	Dynamine	sosthenes	sosthenes
Nymphalidae	Ectima	erycinoides
Nymphalidae	Elzunia	humboldt	humboldt
Nymphalidae	Epiphile	chrysites	chrysites
Nymphalidae	Epiphile	dilecta
Nymphalidae	Epiphile	eriopis	eriopis
Nymphalidae	Eresia	carme
Nymphalidae	Eresia	clio	clio
Nymphalidae	Eresia	datis	manto
Nymphalidae	Eresia	emerantia	emerantia
Nymphalidae	Eresia	ithomioides	poecilina
Nymphalidae	Eresia	levina
Nymphalidae	Eresia	phaedima
Nymphalidae	Eresia	polina	polina
Nymphalidae	Eretris	calisto	calisto
Nymphalidae	Eretris	depresissima
Nymphalidae	Eretris	ocellifera	ocellifera
Nymphalidae	Eretris	oculata
Nymphalidae	Eunica	bechina	bechina
Nymphalidae	Eunica	carias	carias
Nymphalidae	Eunica	evelide	evelide
Nymphalidae	Eunica	norica	norica
Nymphalidae	Euptychia	jesia
Nymphalidae	Euptychoides	griphe
Nymphalidae	Euptychoides	laccine
Nymphalidae	Euptychoides	saturnus
Nymphalidae	Forsterinaria	inornata
Nymphalidae	Forsterinaria	neonympha
Nymphalidae	Forsterinaria	pallida	aurita
Nymphalidae	Forsterinaria	pseudinornata
Nymphalidae	Fountainea	nessus	nessus
Nymphalidae	Fountainea	nobilis	pacifica
Nymphalidae	Gnathotriche	exclamationis	exclamationis
Nymphalidae	Godyris	duillia
Nymphalidae	Godyris	zavaleta
Nymphalidae	Greta	andromica	andromica
Nymphalidae	Hamadryas	fornax
Nymphalidae	Hamadryas	laodamia
Nymphalidae	Heliconius	charithonia	charithonia
Nymphalidae	Heliconius	clysonymus	clysonymus
Nymphalidae	Heliconius	cydno	cydnides
Nymphalidae	Heliconius	doris	obscuris, viridis & dives
Nymphalidae	Heliconius	erato	chestertonii
Nymphalidae	Heliconius	hecale
Nymphalidae	Heliconius	melpomene	martinae
Nymphalidae	Hermeuptychia	cucullina
Nymphalidae	Hermeuptychia	harmonia
Nymphalidae	Hermeuptychia	hermes
Nymphalidae	Hermeuptychia	maimoune
Nymphalidae	Historis	odius	odius
Nymphalidae	Hypanartia	dione	dione
Nymphalidae	Hypanartia	lethe
Nymphalidae	Hypanartia	trimaculata	trimaculata
Nymphalidae	Hypna	clytemnestra
Nymphalidae	Hypoleria	lavinia	vanilia
Nymphalidae	Hypothyris	lycaste
Nymphalidae	Ithomia	avella	cesleria
Nymphalidae	Ithomia	hyala	sincerus
Nymphalidae	Janatella	fellula
Nymphalidae	Janatella	leucodesma
Nymphalidae	Junonia	evarete	zonalis
Nymphalidae	Lycorea	halia
Nymphalidae	Lymanopoda	albocincta	albocincta
Nymphalidae	Lymanopoda	obsoleta
Nymphalidae	Magneuptychia	libye
Nymphalidae	Magneuptychia	tiessa
Nymphalidae	Manataria	hercyna	daguana
Nymphalidae	Marpesia	chiron
Nymphalidae	Marpesia	corinna
Nymphalidae	Marpesia	furcula
Nymphalidae	Marpesia	marcella	marcella
Nymphalidae	Marpesia	petreus
Nymphalidae	Marpesia	zerynthia
Nymphalidae	Mechanitis	polymnia	veritabilis
Nymphalidae	Melinaea	idea
Nymphalidae	Memphis	acidalia	memphis
Nymphalidae	Memphis	arginussa
Nymphalidae	Memphis	laura	caucana
Nymphalidae	Memphis	lyceus
Nymphalidae	Memphis	moeris
Nymphalidae	Memphis	oenomais
Nymphalidae	Memphis	perenna	austrina
Nymphalidae	Memphis	xenocles
Nymphalidae	Mesotaenia	vaninka	doris
Nymphalidae	Morpho	deidamia	granadensis
Nymphalidae	Morpho	helenor	leontius
Nymphalidae	Napeogenes	peridia	hoppi
Nymphalidae	Nessaea	aglaura	aglaura
Nymphalidae	Nica	flavilla
Nymphalidae	Oleria	amalda	amalda
Nymphalidae	Oleria	makrena
Nymphalidae	Oleria	phenomoe
Nymphalidae	Oleria	santineza
Nymphalidae	Opsiphanes	cassina	chiriquensis
Nymphalidae	Opsiphanes	invirae	sticheli
Nymphalidae	Opsiphanes	quiteria	badius
Nymphalidae	Oressinoma	typhla	typhla
Nymphalidae	Oxeoschistus	simplex	fuscus
Nymphalidae	Panacea	prola
Nymphalidae	Parataygetis	lineata	lineata
Nymphalidae	Pareuptychia	hesionides
Nymphalidae	Pareuptychia	metaleuca
Nymphalidae	Pareuptychia	ocirrhoe	ocirrhoe
Nymphalidae	Patricia	dercyllidas	dercyllidas
Nymphalidae	Pedaliodes	dejecta
Nymphalidae	Pedaliodes	demarmelsi
Nymphalidae	Pedaliodes	phrasicla	phrasicla
Nymphalidae	Perisama	bomplandii	bomplandii
Nymphalidae	Perisama	humboldtii	humboldtii
Nymphalidae	Perisama	oppelii	oppelii
Nymphalidae	Perisama	typhania
Nymphalidae	Perisama	yeba	malvina
Nymphalidae	Philaethria	dido	dido
Nymphalidae	Pierella	helvina	helvina
Nymphalidae	Podotricha	judith	caucana
Nymphalidae	Pronophila	unifasciata	brennus
Nymphalidae	Pseudohaetera	hypaesia
Nymphalidae	Pseudohaetera	mimica
Nymphalidae	Pseudomaniola	phaselis	pholoe
Nymphalidae	Pseudoscada	timna	saturata
Nymphalidae	Pyrrhogyra	crameri	undine
Nymphalidae	Pyrrhogyra	edocla	edocla
Nymphalidae	Siproeta	epaphus
Nymphalidae	Siproeta	stelenes
Nymphalidae	Smyrna	blomfildia	datis
Nymphalidae	Taygetis	chrysogone
Nymphalidae	Taygetis	thamyra
Nymphalidae	Tegosa	claudina	claudina
Nymphalidae	Temenis	laothoe	colombiana
Nymphalidae	Tigridia	acesta	acesta
Nymphalidae	Tithorea	tarricina	parola
Nymphalidae	Vanessa	braziliensis
Nymphalidae	Ypthimoides	renata
Papilionidae	Battus	polydamas	polydamas
Papilionidae	Eurytides	colombus
Papilionidae	Heraclides	thoas	nealces
Papilionidae	Neographium	agesilaus	eimeri
Papilionidae	Parides	erithalion	erithalion
Pieridae	Anteos	maerula
Pieridae	Archonias	brassolis	semialba
Pieridae	Ascia	monuste	monuste
Pieridae	Catasticta	ctemene	strigosa
Pieridae	Catasticta	flisa	dilutior
Pieridae	Catasticta	hegemon	helle
Pieridae	Catasticta	prioneris	albescens
Pieridae	Catasticta	sisamnus
Pieridae	Dismorphia	crisia	foedora
Pieridae	Dismorphia	hippotas	diana
Pieridae	Dismorphia	lelex	valeria
Pieridae	Dismorphia	lua	idea
Pieridae	Dismorphia	medora	medora
Pieridae	Dismorphia	mirandola	discolora
Pieridae	Dismorphia	theucharila	theucharila
Pieridae	Dismorphia	zaela	zaela
Pieridae	Dismorphia	zathoe	zathoe
Pieridae	Enantia	lina	agatha
Pieridae	Eurema	albula	marginella
Pieridae	Eurema	arbela	gratiosa
Pieridae	Eurema	daira	lydia
Pieridae	Eurema	elathea	vitellina
Pieridae	Eurema	xantochlora	xantochlora
Pieridae	Hesperocharis	marchallii
Pieridae	Leodonta	tellane	intermedia
Pieridae	Leptophobia	aripa	aripa
Pieridae	Leptophobia	caesia	tenuicornis
Pieridae	Leptophobia	tovaria	maruga
Pieridae	Lieinix	nemesis	nemesis
Pieridae	Melete	leucanthe	leucanthe
Pieridae	Phoebis	neocypris	rurina
Pieridae	Phoebis	philea
Pieridae	Phoebis	sennae
Pieridae	Pseudopieris	nehemia	luisa
Pieridae	Pyrisitia	nise	stygma
Pieridae	Pyrisitia	venusta	venusta
Riodinidae	Amphiselenis	chama
Riodinidae	Ancyluris	inca	huascar
Riodinidae	Ancyluris	jurgensenii	atahualpa
Riodinidae	Anteros	cruentatus
Riodinidae	Anteros	formosus	theleia
Riodinidae	Anteros	kupris	kupris
Riodinidae	Argyrogrammana	caelestina
Riodinidae	Baeotis	macularia
Riodinidae	Baeotis	zonata
Riodinidae	Calephelis	laverna	laverna
Riodinidae	Caria	mantinea
Riodinidae	Caria	rhacotis	rhacotis
Riodinidae	Chamaelimas	briola
Riodinidae	Chalodetta	lypera
Riodinidae	Charis	anius	anius
Riodinidae	Crocozona	pheretima
Riodinidae	Detritivora	hermodora
Riodinidae	Emesis	cypria	paphia
Riodinidae	Emesis	glaucescens
Riodinidae	Emesis	lucinda
Riodinidae	Emesis	lupina
Riodinidae	Emesis	mandana	mandana
Riodinidae	Emesis	ocypore
Riodinidae	Emesis	tegula
Riodinidae	Emesis	tenedia
Riodinidae	Eurybia	lycisca
Riodinidae	Euselasia	bettina
Riodinidae	Euselasia	corduena	anadema
Riodinidae	Euselasia	eupatra
Riodinidae	Exoplisia	hypochalybe
Riodinidae	Lasaia	agesilas	agesilas
Riodinidae	Leucochimona	icare	polita
Riodinidae	Leucochimona	lagora
Riodinidae	Lyropteryx	apollonia
Riodinidae	Melanis	cinaron
Riodinidae	Melanis	electron	pronostriga
Riodinidae	Menander	pretus
Riodinidae	Mesosemia	mevania	mevania
Riodinidae	Mesosemia	minos	modica
Riodinidae	Mesosemia	pacifica
Riodinidae	Mesosemia	zonalis
Riodinidae	Napaea	theages	asteria
Riodinidae	Necyria	bellona	zaneta & westwoodi
Riodinidae	Notheme	erota	diadema
Riodinidae	Rhetus	dysonii	caligosus
Riodinidae	Rhetus	periander	laonome
Riodinidae	Sarota	chrysus
Riodinidae	Sarota	neglecta
Riodinidae	Siseme	alectryo	alectryo
Riodinidae	Siseme	aristoteles	sprucei
Riodinidae	Siseme	pallas	pallas
Riodinidae	Synargis	calyce
Riodinidae	Theope	pedias
Riodinidae	Thisbe	irenea

Butterflies of the Amazon & Andes

Costa Rica July 1994

learnbutterflies.com — Tue, 19 Mar 2024 01:57:38 +0000

This was our second trip to Costa Rica, having organized and led a small group tour there in 1992. The trip was timed to coincide with the Costa Rican rainy season, a time when butterflies emerge en masse in the seasonally dry Guanacaste region in the north. The price to be paid, of course, was that the regions we visited further south were heavily inundated, so butterfly watching in these areas was confined to mornings and early afternoons, before the arrival of the torrential daily rains.

We spent the first 4 days at the American-owned Rancho Naturalista at Tuis, Turrialba. The small rancho is set in mid-elevation transitional rainforest at circa 1400m altitude, with short trails through relict primary forest, secondary forest, and a regenerating clearing.

On sunny mornings, butterflies appeared in excellent numbers with about 95 species recorded. The highlights included Morpho helenor, which patrolled regularly along the tracks, the almost fluorescent orange Dryadula phaetusa, and the stunning black and orange Longwing Heliconius hecale. The most exciting species, however, was an immaculate dazzling iridescent blue Skipper Astraptes fulgerator, which basked in the morning sun for just long enough for one photo. A few moments earlier, we had come across a group of about 30 Phoebis sennae roosting in a small bush. They made a beautiful sight, with the golden morning sunlight streaming through their wings.

At night, the forest was alive with the chirping of myriads of frogs and insects. I left a tungsten lamp switched each night hoping to attract a few moths onto the verandah. Each morning we woke before dawn to investigate and found the walls and ceiling covered in several hundred moths. The variety of shapes, cryptic patterns, and bizarre resting postures adopted by the moths was amazing. There were hordes of beautiful Geometrids, masses of brightly colored Pyralidae, lots of very attractive Tussocks, Prominents, and Leopards, some stunning Hawkmoths, and a couple of Automeris Bull’s-eye Silkmoths. Certainly the greatest moth bonanza in all my years of traveling.

From Rancho Naturalista, we then transferred by road to Los Innocentes, a cattle ranch set below Volcan Orosi in the dry Guanacaste region. For 4 days we were able to explore a variety of habitats including lush grassland, flowery cattle pastures, relict riverbank forest, and a small orchard where fermenting fruit attracted many butterflies. Many beautiful species abounded, including the Crackers Hamadryas feronia, februa, and the incredibly beautiful calico blue amphinome, all perching head-downwards on the tree trunks. Other species included the brilliant orange Dryas iulia nectaring at Lantana, a stunning blue Skipper Paches loxus, the dazzling blue Morpho helenor, and the gorgeous green and brown Malachite Siproeta stelenes.

Along the muddy edges of a stream, a group of Phoebis sennae congregated at cattle urine. Other puddling species included the cream-banded Queen Swallowtail Heraclides androgeus, the gorgeous Red Daggerwing Marpesia petreus, probably the most beautifully shaped butterfly in the world, and the skipper Atarnes sallei, prettily marked in white, orange, and black. In addition to butterflies, Los Innocentes had helicopter damselflies, beautiful cicadas, and several huge and attractive crickets.

Our next destination was Selva Verde, situated on the Rio Sarapiqui, and surrounded by primary and secondary forest, cattle pasture, and banana and palm plantations. We had torrential rain every afternoon commencing about 3 pm, but in the mornings we had about 4 hours to explore the many butterflies in the surrounding forest.

The primary forest behind the lodge was not especially productive, but across the road there were a few hectares of mildly degraded forest where dozens of beautiful species flew along the main track and on the plethora of narrow forest trails. Among the highlights were Morpho helenor, several very attractive Pyrgine skippers, the beautiful transparent Satyrines Cithaerias pireta and Dulcedo polita, and the stunning Metalmark Eurybia lycisca, seen nectaring at Heliconia flowers. We were also very fortunate to get close to a group of Parides childrenae, whose black wings are marked with patches of scarlet and metallic green. To see these lovely butterflies fluttering around bushes at the edge of the forest was certainly one of the highlights of the trip.

We rounded off the trip with a relaxing 3 days at Marenco, a coastal biological station on the Osa peninsula. Butterflies here were generally scarcer than at the other sites but included a few species such as Pierella luna, Euptoieta hegesia, and Heliconius sapho that we had not seen elsewhere.

The behavior of Heliconius erato was fascinating to watch. We found a group of males aggressively jostling around a chrysalis, attempting to copulate with it. A frantic battle took place the instant the female hatched from the chrysalis, as all the males struggled to copulate with her, not even giving her time to expand and dry her wings. One of the males succeeded, and the mated pair then had to endure the attention of the remaining males which were extremely persistent, constantly trying to prise the pair apart. Eventually, as dusk approached, the unsuccessful males started to disperse. I found the pair again the next morning still copulated, with 2 persistent males that had returned and were continuing to harass them.

Species recorded in Costa Rica in July 1994

All species listed below were identified from photographs by Adrian Hoskins. The Papilionoidea were identified using DeVries and d’Abrera for reference. The Hesperioidea were identified using Lewis.

Site	Family	subfamily	Genus	species
Marenco	Hesperiidae	Hesperiinae	Anatrytone	barbara
R. Naturalista	Hesperiidae	Hesperiinae	Anatrytone	perfida
R. Naturalista	Hesperiidae	Hesperiinae	Anthoptus	epictetus
R. Naturalista	Hesperiidae	Hesperiinae	Callimormus	alsimo
R. Naturalista	Hesperiidae	Hesperiinae	Callimormus	corades
Selva Verde	Hesperiidae	Hesperiinae	Calpodes	ethlius
Selva Verde	Hesperiidae	Hesperiinae	Cymaenes	tripunctus
Marenco	Hesperiidae	Hesperiinae	Decinea	percosius
Marenco	Hesperiidae	Hesperiinae	Nastra	julia
Marenco	Hesperiidae	Hesperiinae	Panoquina	lucas
Selva Verde	Hesperiidae	Hesperiinae	Parphorus	storax
R. Naturalista	Hesperiidae	Hesperiinae	Polites	vibex
Selva Verde	Hesperiidae	Hesperiinae	Saliana	placens
R. Naturalista	Hesperiidae	Hesperiinae	Synapte	malitiosa
Marenco	Hesperiidae	Hesperiinae	Vehilius	inca
Selva Verde	Hesperiidae	Hesperiinae	Vehilius	inca
Los Innocentes	Hesperiidae	Pyrginae	Achlyodes	busirus
R. Naturalista	Hesperiidae	Pyrginae	Achlyodes	busirus
Los Innocentes	Hesperiidae	Pyrginae	Achlyodes	mithridates
Selva Verde	Hesperiidae	Pyrginae	Achlyodes	mithridates
Los Innocentes	Hesperiidae	Pyrginae	Aguna	asander
Los Innocentes	Hesperiidae	Pyrginae	Anisochoria	pedaliodina
Los Innocentes	Hesperiidae	Pyrginae	Antigonus	erosus
Los Innocentes	Hesperiidae	Pyrginae	Astraptes	alector
R. Naturalista	Hesperiidae	Pyrginae	Astraptes	fulgerator
Los Innocentes	Hesperiidae	Pyrginae	Atarnes	sallei
Marenco	Hesperiidae	Pyrginae	Autochton	bipunctatus
R. Naturalista	Hesperiidae	Pyrginae	Autochton	neis
R. Naturalista	Hesperiidae	Pyrginae	Autochton	neis
Los Innocentes	Hesperiidae	Pyrginae	Bolla	giselus
Marenco	Hesperiidae	Pyrginae	Chioides	catillus
Los Innocentes	Hesperiidae	Pyrginae	Cogia	cajeta
Selva Verde	Hesperiidae	Pyrginae	Cycloglypha	thrasibulus
Selva Verde	Hesperiidae	Pyrginae	Gorgythion	begga
R. Naturalista	Hesperiidae	Pyrginae	Grais	stigmaticus
Selva Verde	Hesperiidae	Pyrginae	Helias	phalaenoides
Los Innocentes	Hesperiidae	Pyrginae	Heliopetes	arsalte
Marenco	Hesperiidae	Pyrginae	Heliopetes	arsalte
R. Naturalista	Hesperiidae	Pyrginae	Heliopetes	arsalte
Los Innocentes	Hesperiidae	Pyrginae	Heliopetes	laviana
Los Innocentes	Hesperiidae	Pyrginae	Heliopetes	petrus
Los Innocentes	Hesperiidae	Pyrginae	Paches	loxus
Los Innocentes	Hesperiidae	Pyrginae	Polythrix	asine
Los Innocentes	Hesperiidae	Pyrginae	Pyrgus	oileus
Marenco	Hesperiidae	Pyrginae	Pyrgus	oileus
R. Naturalista	Hesperiidae	Pyrginae	Pyrgus	oileus
Selva Verde	Hesperiidae	Pyrginae	Pyrgus	oileus
Selva Verde	Hesperiidae	Pyrginae	Sostrata	bifasciata
Selva Verde	Hesperiidae	Pyrginae	Staphylus	ceos
Selva Verde	Hesperiidae	Pyrginae	Staphylus	mazans
Selva Verde	Hesperiidae	Pyrginae	Telemiades	amphione
R. Naturalista	Hesperiidae	Pyrginae	Urbanus	dorantes
Los Innocentes	Hesperiidae	Pyrginae	Urbanus	doryssus
Los Innocentes	Hesperiidae	Pyrginae	Urbanus	esmeraldus
Los Innocentes	Hesperiidae	Pyrginae	Urbanus	proteus
Marenco	Hesperiidae	Pyrginae	Urbanus	proteus
R. Naturalista	Hesperiidae	Pyrginae	Urbanus	proteus
Marenco	Hesperiidae	Pyrginae	Urbanus	simplicius
R. Naturalista	Hesperiidae	Pyrginae	Urbanus	simplicius
Marenco	Hesperiidae	Pyrginae	Urbanus	teleus
R. Naturalista	Hesperiidae	Pyrginae	Urbanus	teleus
R. Naturalista	Hesperiidae	Pyrginae	Urbanus	viterboana
Los Innocentes	Hesperiidae	Pyrginae	Xenophanes	tryxus
Marenco	Hesperiidae	Pyrginae	Xenophanes	tryxus
Selva Verde	Hesperiidae	Pyrginae	Xenophanes	tryxus
R. Naturalista	Lycaenidae	Polyommatinae	Cupido	comyntas
Los Innocentes	Lycaenidae	Theclinae	Arawacus	aetolus
Marenco	Lycaenidae	Theclinae	Arawacus	aetolus
R. Naturalista	Lycaenidae	Theclinae	Arawacus	aetolus
Selva Verde	Lycaenidae	Theclinae	Arawacus	aetolus
R. Naturalista	Lycaenidae	Theclinae	Arawacus	sito
Los Innocentes	Lycaenidae	Theclinae	Calycopis	isobeon
Marenco	Lycaenidae	Theclinae	Calycopis	isobeon
R. Naturalista	Lycaenidae	Theclinae	Calycopis	isobeon
Marenco	Lycaenidae	Theclinae	Calycopis	pisis
Selva Verde	Lycaenidae	Theclinae	Calycopis	pisis
Selva Verde	Lycaenidae	Theclinae	Calycopis	puppius
Selva Verde	Lycaenidae	Theclinae	Electrostrymon	endymion
R. Naturalista	Lycaenidae	Theclinae	Laothus	barajo
Selva Verde	Lycaenidae	Theclinae	Ostrinotes	halciones
R. Naturalista	Lycaenidae	Theclinae	Rekoa	meton
Marenco	Lycaenidae	Theclinae	Strymon	bazochii
R. Naturalista	Lycaenidae	Theclinae	Strymon	megarus
R. Naturalista	Lycaenidae	Theclinae	Strymon	megarus
Los Innocentes	Lycaenidae	Theclinae	Strymon	mulucha
Marenco	Lycaenidae	Theclinae	Strymon	rufofusca
Selva Verde	Lycaenidae	Theclinae	Strymon	serapio
Los Innocentes	Lycaenidae	Theclinae	Tmolus	echion
Marenco	Lycaenidae	Theclinae	Tmolus	echion
R. Naturalista	Lycaenidae	Theclinae	Tmolus	echion
Los Innocentes	Nymphalidae	Biblidinae	Biblis	hyperia
Los Innocentes	Nymphalidae	Biblidinae	Dynamine	postverta
Los Innocentes	Nymphalidae	Biblidinae	Eunica	mygdonia
Los Innocentes	Nymphalidae	Biblidinae	Hamadryas	amphinome
Los Innocentes	Nymphalidae	Biblidinae	Hamadryas	februa
Los Innocentes	Nymphalidae	Biblidinae	Hamadryas	feronia
Los Innocentes	Nymphalidae	Biblidinae	Hamadryas	glauconome
R. Naturalista	Nymphalidae	Biblidinae	Marpesia	berania
R. Naturalista	Nymphalidae	Biblidinae	Marpesia	chiron
Los Innocentes	Nymphalidae	Biblidinae	Marpesia	petreus
Los Innocentes	Nymphalidae	Biblidinae	Mestra	dorcas
Los Innocentes	Nymphalidae	Biblidinae	Pyrrhogyra	otolais
Los Innocentes	Nymphalidae	Charaxinae	Archaeoprepona	amphimachus
R. Naturalista	Nymphalidae	Charaxinae	Archaeoprepona	amphimachus
Los Innocentes	Nymphalidae	Charaxinae	Archaeoprepona	demophon
R. Naturalista	Nymphalidae	Charaxinae	Archaeoprepona	demophon
R. Naturalista	Nymphalidae	Charaxinae	Fountainea	glycerium
Los Innocentes	Nymphalidae	Charaxinae	Memphis	forreri
Los Innocentes	Nymphalidae	Charaxinae	Prepona	laertes
Selva Verde	Nymphalidae	Charaxinae	Siderone	galanthis
Los Innocentes	Nymphalidae	Danainae	Danaus	eresimus
Los Innocentes	Nymphalidae	Danainae	Danaus	gilippus
Los Innocentes	Nymphalidae	Danainae	Danaus	plexippus
R. Naturalista	Nymphalidae	Danainae	Danaus	plexippus
Selva Verde	Nymphalidae	Danainae	Lycorea	halia
Los Innocentes	Nymphalidae	Heliconiinae	Agraulis	vanillae
Marenco	Nymphalidae	Heliconiinae	Agraulis	vanillae
R. Naturalista	Nymphalidae	Heliconiinae	Dione	juno
Los Innocentes	Nymphalidae	Heliconiinae	Dryadula	phaetusa
R. Naturalista	Nymphalidae	Heliconiinae	Dryadula	phaetusa
Los Innocentes	Nymphalidae	Heliconiinae	Dryas	iulia
Marenco	Nymphalidae	Heliconiinae	Dryas	iulia
R. Naturalista	Nymphalidae	Heliconiinae	Dryas	iulia
Selva Verde	Nymphalidae	Heliconiinae	Dryas	iulia
R. Naturalista	Nymphalidae	Heliconiinae	Eueides	aliphera
Marenco	Nymphalidae	Heliconiinae	Euptoieta	hegesia
Los Innocentes	Nymphalidae	Heliconiinae	Heliconius	charithonia
Marenco	Nymphalidae	Heliconiinae	Heliconius	charithonia
R. Naturalista	Nymphalidae	Heliconiinae	Heliconius	charithonia
Selva Verde	Nymphalidae	Heliconiinae	Heliconius	charithonia
Los Innocentes	Nymphalidae	Heliconiinae	Heliconius	cydno
R. Naturalista	Nymphalidae	Heliconiinae	Heliconius	cydno
Los Innocentes	Nymphalidae	Heliconiinae	Heliconius	erato
Marenco	Nymphalidae	Heliconiinae	Heliconius	erato
R. Naturalista	Nymphalidae	Heliconiinae	Heliconius	erato
Selva Verde	Nymphalidae	Heliconiinae	Heliconius	erato
Los Innocentes	Nymphalidae	Heliconiinae	Heliconius	hecale
Marenco	Nymphalidae	Heliconiinae	Heliconius	hecale
R. Naturalista	Nymphalidae	Heliconiinae	Heliconius	hecale
R. Naturalista	Nymphalidae	Heliconiinae	Heliconius	hecalesia
Selva Verde	Nymphalidae	Heliconiinae	Heliconius	ismenius
Los Innocentes	Nymphalidae	Heliconiinae	Heliconius	melpomene
Marenco	Nymphalidae	Heliconiinae	Heliconius	melpomene
Selva Verde	Nymphalidae	Heliconiinae	Heliconius	melpomene
Marenco	Nymphalidae	Heliconiinae	Heliconius	sapho
Marenco	Nymphalidae	Heliconiinae	Heliconius	sara
R. Naturalista	Nymphalidae	Heliconiinae	Heliconius	sara
Selva Verde	Nymphalidae	Heliconiinae	Heliconius	sara
Marenco	Nymphalidae	Heliconiinae	Laparus	doris
R. Naturalista	Nymphalidae	Heliconiinae	Laparus	doris
Selva Verde	Nymphalidae	Heliconiinae	Laparus	doris
Los Innocentes	Nymphalidae	Heliconiinae	Philaethria	dido
Marenco	Nymphalidae	Heliconiinae	Philaethria	dido
R. Naturalista	Nymphalidae	Ithomiinae	Godyris	nero
Marenco	Nymphalidae	Ithomiinae	Ithomia	patilla
R. Naturalista	Nymphalidae	Ithomiinae	Ithomia	patilla
Los Innocentes	Nymphalidae	Ithomiinae	Mechanitis	lysimnia
R. Naturalista	Nymphalidae	Ithomiinae	Mechanitis	lysimnia
Selva Verde	Nymphalidae	Ithomiinae	Mechanitis	lysimnia
Los Innocentes	Nymphalidae	Ithomiinae	Mechanitis	menapis
R. Naturalista	Nymphalidae	Ithomiinae	Mechanitis	menapis
Los Innocentes	Nymphalidae	Ithomiinae	Mechanitis	polymnia
Los Innocentes	Nymphalidae	Ithomiinae	Melinaea	lilis
Selva Verde	Nymphalidae	Ithomiinae	Melinaea	lilis
R. Naturalista	Nymphalidae	Ithomiinae	Tithorea	tarricina
R. Naturalista	Nymphalidae	Limenitidinae	Adelpha	barnesia
Los Innocentes	Nymphalidae	Limenitidinae	Adelpha	cytherea
Marenco	Nymphalidae	Limenitidinae	Adelpha	cytherea
R. Naturalista	Nymphalidae	Limenitidinae	Adelpha	cytherea
Selva Verde	Nymphalidae	Limenitidinae	Adelpha	cytherea
R. Naturalista	Nymphalidae	Limenitidinae	Adelpha	serpa
R. Naturalista	Nymphalidae	Limenitidinae	Adelpha	zina
R. Naturalista	Nymphalidae	Melitaeinae	Microtia	elva
Selva Verde	Nymphalidae	Morphinae	Antirrhea	philoctetes
Selva Verde	Nymphalidae	Morphinae	Antirrhea	philoctetes
R. Naturalista	Nymphalidae	Morphinae	Antirrhea	pterocopha
Selva Verde	Nymphalidae	Morphinae	Caligo	atreus
Selva Verde	Nymphalidae	Morphinae	Caligo	brasiliensis
Los Innocentes	Nymphalidae	Morphinae	Caligo	telamonius
Selva Verde	Nymphalidae	Morphinae	Eryphanis	automedon
Los Innocentes	Nymphalidae	Morphinae	Mimoblepia	staudingeri
Los Innocentes	Nymphalidae	Morphinae	Morpho	helenor
Marenco	Nymphalidae	Morphinae	Morpho	helenor
R. Naturalista	Nymphalidae	Morphinae	Morpho	helenor
Selva Verde	Nymphalidae	Morphinae	Morpho	helenor
Selva Verde	Nymphalidae	Morphinae	Opsiphanes	cassina
Los Innocentes	Nymphalidae	Morphinae	Opsiphanes	tamarindi
Los Innocentes	Nymphalidae	Nymphalinae	Anartia	fatima
Marenco	Nymphalidae	Nymphalinae	Anartia	fatima
R. Naturalista	Nymphalidae	Nymphalinae	Anartia	fatima
Selva Verde	Nymphalidae	Nymphalinae	Anartia	fatima
Los Innocentes	Nymphalidae	Nymphalinae	Anartia	jatrophae
Marenco	Nymphalidae	Nymphalinae	Anartia	jatrophae
R. Naturalista	Nymphalidae	Nymphalinae	Anartia	jatrophae
Selva Verde	Nymphalidae	Nymphalinae	Anartia	jatrophae
Marenco	Nymphalidae	Nymphalinae	Anthanassa	ardys
Los Innocentes	Nymphalidae	Nymphalinae	Anthanassa	drusilla
Los Innocentes	Nymphalidae	Nymphalinae	Anthanassa	frisia
Los Innocentes	Nymphalidae	Nymphalinae	Chlosyne	erodyle